Thin-walled spores produced by the uredium
Urediniospores of 11 Milesina species. a Milesina blechni on Struthiopteris spicant b Milesina blechni on Struthiopteris spicant , cracked spore with released plasma, germ pores scattered c Milesina carpatica on Dryopteris filix-mas d Milesina exigua on Polystichum braunii , smooth surface e Milesina exigua on Polystichum braunii , smooth surface, plasma-free spore, germ pores bipolar f Milesina feurichii on Asplenium septentrionale with smooth areas on surface g Milesina feurichii on Asplenium septentrionale , cracked plasma-free spore, germ pores scattered h Milesina kriegeriana on Dryopteris carthusiana i Milesina magnusiana on Asplenium adiantum-nigrum with smooth areas on surface j Milesina magnusiana on Asplenium adiantum-nigrum , spore plasma-free, germ pores scattered k Milesina murariae on Asplenium ruta-muraria with smooth areas on surface l Milesina murariae on Asplenium ruta-muraria , cracked spore with released plasma, germ pores scattered m Milesina polypodii on Polypodium vulgare with smooth areas on surface n Milesina scolopendrii on Asplenium scolopendrium with smooth areas on surface o Milesina vogesiaca on Polystichum aculeatum , surface with very flat warts at the tip of the spore (arrow) p Milesina vogesiaca on Polystichum aculeatum , surface smooth (no warts visible at the tip), germ pores bipolar q Milesina whitei on Polystichum sp. r Milesina whitei on Polystichum setiferum , cracked spore with released plasma, germ pores scattered. Urediniospores (or uredospores ) are thin-walled spores produced by the uredium , a stage in the life-cycle of rusts .
Development
Urediniospores develop in the uredium , generally on a leaf's under surface.