Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Acanthophyllum filiforme^[8]	Sp. nov	Valid	Coen-Aubert	Devonian (Givetian)		Mauritania	A rugose coral belonging to the family Ptenophyllidae.
Acanthophyllum sougyi^[8]	Sp. nov	Valid	Coen-Aubert	Devonian (Givetian)		Mauritania	A rugose coral belonging to the family Ptenophyllidae.
Agetolites angullongensis^[9]	Sp. nov	Valid	Zhen, Wang & Percival	Late Ordovician	Angullong Formation	Australia	A tabulate coral.
Aulohelia carbonica^[10]	Sp. nov	Valid	Niko & Fujikawa	Carboniferous (Viséan)	Akiyoshi Limestone Group	Japan	A tabulate coral.
Bothrophyllum gorbachevensis^[11]	Sp. nov	Valid	Fedorowski	Carboniferous (Bashkirian)		Ukraine	A rugose coral belonging to the family Bothrophyllidae.
Bothrophyllum kalmyussi^[11]	Sp. nov	Valid	Fedorowski	Carboniferous (Bashkirian)		Ukraine	A rugose coral belonging to the family Bothrophyllidae.
Cambroctoconus koori^[12]	Sp. nov	Valid	Peel	Cambrian Stage 4 or Stage 5	Henson Gletscher Formation	Greenland	A possible member of Octocorallia.
Charactophyllum mauritanicum^[8]	Sp. nov	Valid	Coen-Aubert	Devonian (Givetian)		Mauritania	A rugose coral belonging to the family Disphyllidae.
Charactophyllum soraufi^[8]	Sp. nov	Valid	Coen-Aubert	Devonian (Givetian)		Mauritania	A rugose coral belonging to the family Disphyllidae.
Dianqianophyllum^[13]	Gen. et sp. nov	Valid	Liao & Ma	Devonian (Givetian)		China	A rugose coral. Genus includes new species D. bianqingense.
Dibunophylloides columnatus^[14]	Sp. nov	Valid	Fedorowski	Carboniferous (Bashkirian)		Ukraine	A rugose coral belonging to the family Aulophyllidae.
Dibunophylloides paulus^[14]	Sp. nov	Valid	Fedorowski	Carboniferous (Bashkirian)		Ukraine	A rugose coral belonging to the family Aulophyllidae.
Dibunophylloides similis^[14]	Sp. nov	Valid	Fedorowski	Carboniferous (Bashkirian)		Ukraine	A rugose coral belonging to the family Aulophyllidae.
Dibunophyllum medium^[14]	Sp. nov	Valid	Fedorowski	Carboniferous (Bashkirian)		Ukraine	A rugose coral belonging to the family Aulophyllidae.
Enniskillenia multiseptata^[15]	Sp. nov	Valid	Bamber & Rodríguez in Bamber et al.	Carboniferous (Mississippian)		Canada	A rugose coral.
Fungiaphyllia^[16]	Gen. et sp. nov	Valid	Melnikova & Roniewicz	Early Jurassic (Hettangian/Sinemurian–Pliensbachian)		Afghanistan	A stony coral belonging to the family Latomeandridae. The type species is Fungiaphyllia communis.
Gillismilia^[17]	Nom. nov	Valid	Lathuilière, Charbonnier & Pacaud	Early Jurassic (Pliensbachian)		France	A coral; a replacement name for Palaeocyathus Alloiteau (1956).
Guembelastraea dronovi^[16]	Sp. nov	Valid	Melnikova & Roniewicz	Early Jurassic (Hettangian/Sinemurian)		Afghanistan	A stony coral belonging to the family Tropiastraeidae, a species of Guembelastraea.
Lithostrotion termieri^[18]	Sp. nov	Valid	Rodríguez & Somerville in Rodríguez, Somerville & Said	Carboniferous (Viséan)	Azrou-Khenifra Basin	Morocco	A rugose coral belonging to the family Lithostrotionidae.
Macgeea tourneuri^[8]	Sp. nov	Valid	Coen-Aubert	Devonian (Givetian)		Mauritania	A rugose coral belonging to the family Phillipsastreidae.
Nina^[11]	Gen. et 3 sp. et comb. nov	Junior homonym	Fedorowski	Carboniferous (Serpukhovian and Bashkirian)		Ukraine	A rugose coral belonging to the family Bothrophyllidae. The type species is N. donetsiana; genus also includes new species N. dibimitaria and N. magna, as well as "Bothrophyllum" berestovensis Vassilyuk (1960). The generic name is preoccupied by Nina Horsfield (1829).
Oppelismilia spectabilis^[16]	Sp. nov	Valid	Melnikova & Roniewicz	Early Jurassic (Hettangian/Sinemurian)		Afghanistan	A stony coral belonging to the family Oppelismiliidae, a species of Oppelismilia.
Parepismilia dolichostoma^[16]	Sp. nov	Valid	Melnikova & Roniewicz	Early Jurassic (Hettangian–early Sinemurian)		Afghanistan	A stony coral belonging to the family Parepismiliidae, a species of Parepismilia.
Parepismilia dronovi^[16]	Sp. nov	Valid	Melnikova & Roniewicz	Early Jurassic (Hettangian/Sinemurian)		Afghanistan	A stony coral belonging to the family Parepismiliidae, a species of Parepismilia.
Periplacotrochus^[19]	Gen. et comb. et sp. nov	Valid	Cairns	Late Eocene to middle Miocene		Australia	A flabellid coral. Genus includes P. deltoideus (Duncan, 1864), P. corniculatus (Dennant, 1899), P. elongatus (Duncan, 1864), P. pueblensis (Dennant, 1903), P. inflectus (Dennant, 1903) and P. magnus (Dennant, 1904), as well as new species P. cudmorei.
Petrophyllia niimiensis^[20]	Sp. nov	Valid	Niko, Suzuki & Taguchi	Miocene	Bihoku Group	Japan	A stony coral.
Protomichelinia funafusensis^[21]	Sp. nov	Valid	Niko	Early Permian	Funafuseyama Limestone	Japan	A tabulate coral belonging to the order Favositida and the family Micheliniidae.
Qinscyphus^[22]	Gen. et sp. nov	Valid	Liu et al.	Cambrian (Fortunian)	Kuanchuanpu Formation	China	A probable crown jellyfish belonging to the family Olivooidae. The type species is Q. necopinus.
Rozkowskia lenta^[14]	Sp. nov	Valid	Fedorowski	Carboniferous (Bashkirian)		Ukraine	A rugose coral belonging to the family Aulophyllidae.
Scoliopora hosakai^[23]	Sp. nov	Valid	Niko, Ibaraki & Tazawa	Middle Devonian		Japan	A tabulate coral belonging to the order Favositida and the family Alveolitidae.
Sinaster^[24]	Gen. et sp. nov	Valid	Wang et al.	Early Cambrian	Kuanchuanpu Formation	China	A member of Medusozoa belonging to the family Olivooidae. The type species is S. petalon.
Stephanophyllia plattenwaldensis^[25]	Sp. nov	Valid	Baron-Szabo	Early Cretaceous (late Aptian to Albian)	Garschella Formation	Austria	A stony coral belonging to the family Micrabaciidae.
Sterictopathes^[26]	Gen. et sp. nov	Valid	Baliński & Sun	Ordovician (early Floian)	Fenxiang Formation Honghuayuan Formation	China	A black coral related to Sinopathes reptans. The type species is S. radicatus.
Voragoaxum^[14]	Gen. et sp. nov	Valid	Fedorowski	Carboniferous (Bashkirian)		Ukraine	A rugose coral belonging to the family Aulophyllidae. The type species is V. cavum.
Zaphrentites etheringtonensis^[15]	Sp. nov	Valid	Bamber & Rodríguez in Bamber et al.	Carboniferous (Mississippian)		Canada	A rugose coral.
Zaphrentites lerandi^[15]	Sp. nov	Valid	Bamber & Rodríguez in Bamber et al.	Carboniferous (Mississippian)		Canada	A rugose coral.

Bryozoans

Research

Epizoic bryozoans are reported on fossil crabs from the Miocene Mishan Formation (Iran) by Key et al. (2017).^[27]

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Acupipora mexicana^[28]	Sp. nov	Valid	Ernst & Vachard	Carboniferous (middle Pennsylvanian)		Mexico
Adeonellopsis sandbergi^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Adeonidae.
'Akatopora' wilmseni^[30]	Sp. nov	Valid	Martha, Niebuhr & Scholz	Late Cretaceous (mid-late Turonian)	Strehlen Formation	Germany	A cheilostome bryozoan.
Atactotoechus vaulxensis^[31]	Sp. nov	Valid	Ernst et al.	Carboniferous (Mississippian)		Belgium	A bryozoan.
Bashkirella arnaoense^[32]	Sp. nov	Valid	Suárez Andrés & Wyse Jackson	Devonian (Eifelian)	Moniello Formation	Spain	A member of Fenestrata belonging to the family Chasmatoporidae.
Bigeyina cantabrica^[32]	Sp. nov	Valid	Suárez Andrés & Wyse Jackson	Devonian (Emsian–early Eifelian)	Moniello Formation	Spain	A member of Fenestrata belonging to the family Semicosciniidae.
Bigeyina spinosa^[32]	Sp. nov	Valid	Suárez Andrés & Wyse Jackson	Devonian (Emsian–early Eifelian)	Moniello Formation	Spain	A member of Fenestrata belonging to the family Semicosciniidae.
Bragella^[33]	Gen. et sp. nov	Valid	Di Martino et al.	Eocene–Oligocene transition		Tanzania	A cheilostome bryozoan. Genus includes new species B. pseudofedora.
Buskia waiinuensis^[34]	Sp. nov	Valid	Di Martino et al.	Pleistocene	Nukumaru Limestone	New Zealand	A member of Ctenostomatida belonging to the superfamily Vesicularioidea and the family Buskiidae.
Cheiloporina clarksvillensis^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Cheiloporinidae.
Cigclisula solenoides^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Colatooeciidae.
Coeloclemis zefrehensis^[35]	Sp. nov	Valid	Ernst et al.	Devonian (Frasnian)	Bahram Formation	Iran	A trepostome bryozoan.
Diplosolen akatjevense^[36]	Sp. nov	Valid	Viskova & Pakhnevich	Middle Jurassic (Callovian)		Russia	A bryozoan belonging to the class Stenolaemata and the order Tubuliporida.
Ditaxipora lakriensis^[37]	Sp. nov	Valid	Sonar & Pawar	Miocene (Burdigalian)	Chhasra Formation	India	A member of the family Catenicellidae.
Eridopora moravica^[38]	Sp. nov	Valid	Tolokonnikova, Kalvoda & Kumpan	Carboniferous (Tournaisian)		Czech Republic
Escharoides joannae^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Romancheinidae.
Euthyrhombopora tenuis^[35]	Sp. nov	Valid	Ernst et al.	Devonian (Frasnian)	Bahram Formation	Iran	A rhabdomesine cryptostome bryozoan.
Exechonella minutiperforata^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Exechonellidae.
Exidmonea baghi^[39]	Sp. nov	Valid	Zágoršek, Yazdi & Bahrami	Miocene	Qom Formation	Iran	A cyclostome bryozoan.
Fabifenestella almazani^[28]	Sp. nov	Valid	Ernst & Vachard	Carboniferous (middle Pennsylvanian)		Mexico
Fenestrapora elegans^[32]	Sp. nov	Valid	Suárez Andrés & Wyse Jackson	Devonian (late Emsian–early Eifelian)	Moniello Formation	Spain	A member of Fenestrata belonging to the family Semicosciniidae.
Filites robustus^[32]	Sp. nov	Valid	Suárez Andrés & Wyse Jackson	Devonian (Emsian–early Eifelian)	Moniello Formation	Spain	A member of Fenestrata belonging to the family Acanthocladiidae.
Floridina subantiqua^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Onychocellidae.
Foratella cervisia^[40]	Sp. nov	Valid	Taylor & Martha	Late Cretaceous (Cenomanian)	Beer Head Limestone Formation	United Kingdom	A cheilostome bryozoan.
Hagiosynodos simplex^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Cheiloporinidae.
Heteractis tanzaniensis^[33]	Sp. nov	Valid	Di Martino et al.	Eocene–Oligocene transition		Tanzania	A cheilostome bryozoan.
Hillmeropora^[30]	Gen. et sp. nov	Valid	Martha, Niebuhr & Scholz	Late Cretaceous (mid-late Turonian)	Strehlen Formation	Germany	A cheilostome bryozoan genus belonging to the family Calloporidae. Type species H. pavonina; genus also includes Membranipora procurrens Brydone, 1929.
Jablonskipora^[41]	Gen. et sp. nov	Valid	Martha & Taylor	Early Cretaceous (Albian)	Upper Greensand	United Kingdom	A cheilostome bryozoan. The type species is J. kidwellae.
Kalvariella antiqua^[32]	Sp. nov	Valid	Suárez Andrés & Wyse Jackson	Devonian (Emsian–early Eifelian)	Moniello Formation	Spain	A member of Fenestrata belonging to the family Acanthocladiidae.
Lacrimula crassa^[33]	Sp. nov	Valid	Di Martino et al.	Eocene–Oligocene transition		Tanzania	A cheilostome bryozoan.
Lacrimula kilwaensis^[33]	Sp. nov	Valid	Di Martino et al.	Eocene–Oligocene transition		Tanzania	A cheilostome bryozoan.
Margaretta pentaceratops^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Margarettidae.
Metrarabdotos aquaeguttum^[42]	Sp. nov	Valid	Ramalho, Távora & Zagorsek	Early Miocene	Pirabas Formation	Brazil	A member of Lepralielloidea belonging to the family Metrarabdotosidae.
Metrarabdotos capanemensis^[42]	Sp. nov	Valid	Ramalho, Távora & Zagorsek	Early Miocene	Pirabas Formation	Brazil	A member of Lepralielloidea belonging to the family Metrarabdotosidae.
Metrarabdotos elongatum^[42]	Sp. nov	Valid	Ramalho, Távora & Zagorsek	Early Miocene	Pirabas Formation	Brazil	A member of Lepralielloidea belonging to the family Metrarabdotosidae.
Microeciella kolomnensis^[36]	Sp. nov	Valid	Viskova & Pakhnevich	Middle Jurassic (Callovian)		Russia	A bryozoan belonging to the suborder Tubuliporina and the family Oncousoeciidae.
Microporella rusti^[34]	Sp. nov	Valid	Di Martino et al.	Pleistocene	Nukumaru Limestone	New Zealand	A member of the family Microporellidae.
Nellia winstonae^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Quadricellariidae.
Nevianipora isfahani^[39]	Sp. nov	Valid	Zágoršek, Yazdi & Bahrami	Miocene	Qom Formation	Iran	A cyclostome bryozoan.
'Onychocella' barbata^[30]	Sp. nov	Valid	Martha, Niebuhr & Scholz	Late Cretaceous (late Cenomanian)	Dölzschen Formation	Germany	A cheilostome bryozoan. Taylor, Martha & Gordon (2018) transferred this species to the genus Kamilocella.^[43]
Onychocella saxoniae^[30]	Sp. nov	Valid	Martha, Niebuhr & Scholz	Late Cretaceous (late Cenomanian)	Dölzschen Formation	Germany	A cheilostome bryozoan.
Paralicornia interdigitata^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Candidae.
Paraseptopora geometrica^[32]	Sp. nov	Valid	Suárez Andrés & Wyse Jackson	Devonian (late Emsian–early Eifelian)	Moniello Formation	Spain	A member of Fenestrata belonging to the family Septoporidae.
Paraseptopora irregularis^[32]	Sp. nov	Valid	Suárez Andrés & Wyse Jackson	Devonian (Emsian–early Eifelian)	Moniello Formation	Spain	A member of Fenestrata belonging to the family Septoporidae.
Pharopora^[44]	Gen. et sp. nov	Valid	Wyse Jackson, Ernst & Suárez Andrés	Carboniferous (Tournaisian)	Hook Head Formation	Ireland	A member of Cryptostomata belonging to the family Rhabdomesidae. The type species is P. regularis.
Pleuromucrum epifanioi^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Phidoloporidae.
Pleuromucrum liowae^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Phidoloporidae.
Polyascosoecia iranica^[39]	Sp. nov	Valid	Zágoršek, Yazdi & Bahrami	Miocene	Qom Formation	Iran	A cyclostome bryozoan.
Puellina quadrispinosa^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Cribrilinidae.
Revalotrypa inopinata^[45]	Sp. nov	Valid	Fedorov, Koromyslova & Martha	Ordovician (Floian)		Russia	An esthonioporate bryozoan belonging to the family Revalotrypidae.
Revalotrypa yugaensis^[45]	Sp. nov	Valid	Fedorov, Koromyslova & Martha	Ordovician (Floian)		Russia	An esthonioporate bryozoan belonging to the family Revalotrypidae.
Schizolepraliella^[29]	Gen. et sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A Schizoporella-like cheilostome bryozoan of uncertain phylogenetic placement. The type species is S. nancyae.
Selenaria lyrulata^[46]	Sp. nov	Valid	López-Gappa, Pérez & Griffin	Early Miocene	Monte León Formation	Argentina	A bryozoan belonging to the family Selenariidae.
Spiniflabellum jacksoni^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Cribrilinidae.
Steginoporella tiara^[47]	Sp. nov	Valid	Gordon, Voje & Taylor	Early Pleistocene		New Zealand	A member of Cheilostomata belonging to the family Steginoporellidae.
Stylopoma farleyensis^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Schizoporellidae.
Stylopoma leverhulme^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Schizoporellidae.
Thalamoporella bitorquata^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Thalamoporellidae.
Thalamoporella hastigera^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Thalamoporellidae.
Thalamoporella ogivalis^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Thalamoporellidae.
Thalamoporella papalis^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Thalamoporellidae.
Thalamoporella polygonalis^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Thalamoporellidae.
Trypostega vokesi^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Trypostegidae.
Turbicellepora giardinai^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Celleporidae.
Utropora parva^[32]	Sp. nov	Valid	Suárez Andrés & Wyse Jackson	Devonian (Emsian–early Eifelian)	Moniello Formation	Spain	A member of Fenestrata belonging to the family Semicosciniidae.
Vix scolaroi^[29]	Sp. nov	Valid	Di Martino, Taylor & Portell	Early Miocene	Chipola Formation	United States ( Florida)	A cheilostome bryozoan belonging to the family Vicidae.
Wilbertopora manubriformis^[40]	Sp. nov	Valid	Taylor & Martha	Late Cretaceous (Cenomanian)	Beer Head Limestone Formation	United Kingdom	A cheilostome bryozoan.
Wilbertopora ostiolatoides^[30]	Sp. nov	Valid	Martha, Niebuhr & Scholz	Late Cretaceous (mid-late Turonian)	Strehlen Formation	Germany	A cheilostome bryozoan.

Brachiopods

Research

A study on the selectivity of brachiopod extinctions during the Ordovician–Silurian extinction events is published by Finnegan, Rasmussen & Harper (2017).^[48]
A study on the patterns of biomineralization of Late Permian brachiopod shells and on their implications for inferring the environmental disruptions associated with the Permian–Triassic extinction event is published by Garbelli, Angiolini & Shen (2017).^[49]

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Acrothyra bonnia^[50]	Sp. nov	Valid	Skovsted et al.	Cambrian Stage 4	Forteau Formation	Canada ( Newfoundland and Labrador)	A member of Acrotretida belonging to the family Acrotretidae.
Anarhynchia smithi^[51]	Sp. nov	Valid	Pálfy et al.	Early Jurassic (Pliensbachian)	Inklin Formation	Canada ( British Columbia)
Atelelasma longisulcum^[52]	Sp. nov	Valid	Liljeroth et al.	Ordovician	Dunabrattin Limestone Formation Tramore Limestone Formation	Ireland	A member of Strophomenata belonging to the order Billingsellida and the family Clitambonitidae.
Atychorhynchia^[53]	Gen. et sp. nov	Valid	Baeza-Carratalá, Reolid & García Joral	Early Jurassic (late Pliensbachian–early Toarcian)	Zegrí Formation	Spain	A member of Rhynchonellida belonging to the family Norellidae. The type species is A. falsiorigo.
Avdeevella^[54]	Gen. et sp. nov	Valid	Baranov	Ordovician		Russia	The type species is A. mica.
Bilobia alichovae^[55]	Sp. nov	Valid	Madison	Ordovician (Sandbian)		Russia ( Leningrad Oblast)	A member of Strophomenida.
Bittnerithyris^[56]	Gen. nov	Valid	Popov & Zakharov	Early Triassic (Olenekian)		Russia ( Primorsky Krai)	A member of Terebratulida.
Bronnothyris danaperensis^[57]	Sp. nov	Valid	Bitner & Müller	Eocene (Priabonian)		Ukraine	A member of Terebratulida belonging to the family Megathyrididae.
Burrirhynchia albiensis^[58]	Sp. nov	Valid	Gaspard	Early Cretaceous (Albian)		France	A member of Rhynchonellida belonging to the family Tetrarhynchiidae.
Colaptomena auduni^[52]	Sp. nov	Valid	Liljeroth et al.	Ordovician	Tramore Limestone Formation	Ireland	A member of Strophomenida belonging to the family Rafinesquinidae.
Cyrtinaella? houi^[59]	Sp. nov	Valid	Lü & Ma	Devonian (late Frasnian)		China	A member of Spiriferinida.
Cyrtospirifer ainosawensis^[60]	Sp. nov	Valid	Tazawa, Inose & Kaneko	Late Devonian	Ainosawa Formation	Japan	A member of Spiriferida belonging to the family Cyrtospiriferidae.
Cyrtospirifer choanjiensis^[61]	Sp. nov	Valid	Tazawa	Late Devonian		Japan	A member of Spiriferida belonging to the family Cyrtospiriferidae.
Dactylogonia costellata^[52]	Sp. nov	Valid	Liljeroth et al.	Ordovician	Dunabrattin Limestone Formation Tramore Limestone Formation	Ireland	A member of Strophomenida belonging to the family Strophomenidae.
Dirafinesquina antiqua^[62]	Sp. nov	Valid	Popov & Cocks	Ordovician (Dapingian)		Iran	A strophomenoid brachiopod.
Discinisca suborbicularis^[63]	Sp. nov	Valid	Smirnova et al.	Late Jurassic		Russia
Discinisca undata^[64]	Sp. nov	Valid	Smirnova in Smirnova et al.	Late Jurassic		Russia	A brachiopod belonging to the family Discinidae, a species of Discinisca.
Elkanathyris^[65]	Gen. et sp. nov	Valid	Copper & Jin	Silurian (Aeronian)		Canada ( Quebec)	An athyride brachiopod. The type species is E. pallula.
Eoporambonites raziabadensis^[62]	Sp. nov	Valid	Popov & Cocks	Ordovician (Dapingian)		Iran	A porambonitoid brachiopod.
Foveola ivari^[66]	Sp. nov	Valid^[67]	Holmer et al.	Ordovician (Sandbian)		Estonia	A member of Obolidae.
Gypidula xui^[59]	Sp. nov	Valid	Lü & Ma	Devonian (late Frasnian)		China	A member of Pentamerida.
Hesperorthis leinsterensis^[52]	Sp. nov	Valid	Liljeroth et al.	Ordovician	Dunabrattin Limestone Formation Tramore Limestone Formation	Ireland	A member of Orthida belonging to the family Hesperorthidae.
Hexigtenichonetes^[68]	Gen. et comb. nov	Valid	Shen in Shen et al.	Permian (Guadalupian)	Miaoling Formation	China	A member of Productida belonging to the family Rugosochonetidae. The type species is "Hemichonetes" hemipleura Li & Su in Li et al. (1980); genus also includes "Hemichonetes guangxingensis Li & Su in Li et al. (1980), "Hemichonetes subquadrata Li & Su in Li et al. (1980) and "Hemichonetes yanjiensis Li & Su in Li et al. (1980).
Hibernobonites^[52]	Gen. et comb. nov	Valid	Liljeroth et al.	Ordovician	Dunabrattin Limestone Formation Tramore Limestone Formation Tourmakeady Limestone Formation?	Ireland	A member of Pentamerida belonging to the family Porambonitidae. The type species is "Atrypa" filosa M'Coy (1846); genus might also include "Porambonites" dubius Williams & Curry (1985).
Howellites hibernicus^[52]	Sp. nov	Valid	Liljeroth et al.	Ordovician	Dunabrattin Limestone Formation Tramore Limestone Formation	Ireland	A member of Orthida belonging to the family Dalmanellidae.
Isophragma parallelum^[52]	Sp. nov	Valid	Liljeroth et al.	Ordovician	Dunabrattin Limestone Formation Tramore Limestone Formation	Ireland	A member of Strophomenida belonging to the family Plectambonitidae.
Joania ukrainica^[57]	Sp. nov	Valid	Bitner & Müller	Eocene (Priabonian)		Ukraine	A member of Terebratulida belonging to the family Megathyrididae.
Karadagithyris boullierae^[69]	Sp. nov	Valid	Halamski & Cherif	Late Jurassic (Oxfordian)	Argiles de Saïda Formation	Algeria	A member of Terebratulida belonging to the family Muirwoodellidae.
Karlsorus^[70]	Gen. et comb. nov	Valid	Jin & Holmer	Silurian (Wenlock)		Sweden	A new genus for "Pentamerus" gothlandicus Lebedev (1892).
Koninckodonta sumuntanensis^[53]	Sp. nov	Valid	Baeza-Carratalá, Reolid & García Joral	Early Jurassic (late Pliensbachian–early Toarcian)	Zegrí Formation	Spain	A member of Athyridida belonging to the family Koninckinidae.
Kurtothyris^[68]	Nom. nov	Valid	Shen in Shen et al.	Permian (late Cisuralian)	Chihsia Formation	China	A member of Spiriferida belonging to the family Skelidorygmidae; a replacement name for Litothyris Chang (1987). The type species is "Litothyris" anhuiensis Chang (1987).
Kyrshabaktella diabola^[50]	Sp. nov	Valid	Skovsted et al.	Cambrian Stage 4	Forteau Formation	Canada ( Newfoundland and Labrador)	A member of Linguloidea belonging to the family Kyrshabaktellidae.
Lacunites ivantsovi^[66]	Sp. nov	Valid^[67]	Holmer et al.	Ordovician (early Darriwilian)		Russia	A paterinid brachiopod.
Lamellaerhynchia carronensis^[58]	Sp. nov	Valid	Gaspard	Early Cretaceous (Albian)		France	A member of Rhynchonellida belonging to the family Cyclothyrididae.
Leptagonia franca^[71]	Sp. nov	Valid	Mottequin & Simon	Carboniferous (Tournaisian)	Tournai Formation	Belgium	A member of Strophomenoidea belonging to the family Rafinesquinidae.
Levipugnax? liui^[59]	Sp. nov	Valid	Lü & Ma	Devonian (late Frasnian)		China	A member of Rhynchonellida.
Liaotarimella^[68]	Nom. nov	Valid	Shen in Shen et al.	Permian (Artinskian)	Wutankule Formation	China	A member of Productida belonging to the family Productellidae. A replacement name for Tarimella Chen (2004). The type species is "Tarimella" tarimensis Chen (2004).
Lichuanorelloides^[72]	Gen. et sp. nov	Valid	Wang et al.	Early Triassic		China	Genus includes new species L. lichuanensis.
Meristella? aksuensis^[73]	Sp. nov	Valid	Modzalevskaya et al.	Devonian (Lochkovian)		Tajikistan
Nisusia guizhouensis^[74]	Sp. nov	Valid	Mao et al.	Cambrian	Kaili Formation Qingxudong Formation	China	A brachiopod belonging to the subphylum Rhynchonelliformea, order Kutorginida and the family Nisusiidae.
Nucleospira hannoniae^[71]	Nom. nov	Valid	Mottequin & Simon	Carboniferous (Tournaisian)	Tournai Formation	Belgium	A member of Athyridida belonging to the family Nucleospiridae; a replacement name for Athyris globulina de Koninck (1887).
Onniella variabilis^[75]	Sp. nov	Valid	Harper, Parkes & Zhan	Ordovician (Katian)	Raheen Formation	Ireland	A dalmanelloid brachiopod belonging to the family Dalmanellidae.
Ouraniorhynchus^[73]	Gen. et sp. nov	Valid	Modzalevskaya et al.	Devonian (Lochkovian)		Tajikistan	A brachiopod. Genus includes new species O. dronovi.
Permocryptospirifer^[68]	Gen. et comb. nov	Valid	Shen & Grunt in Shen et al.	Permian (late Cisuralian and Guadalupian)	Chihsia Formation Maokou Formation Shazipo Formation	China	A member of Athyridida belonging to the family Athyrididae. The type species is "Cryptospirifer" omeishanensis Huang (1933); genus also includes "Cryptospirifer" minor Yang (1984) and "Cryptospirifer" shawanensis Jin et al. (1974).
Piarorhynchella tazawai^[56]	Sp. nov	Valid	Popov & Zakharov	Early Triassic (Olenekian)		Russia ( Primorsky Krai)	A member of Rhynchonellida.
Platystrophia tramorensis^[52]	Sp. nov	Valid	Liljeroth et al.	Ordovician	Tramore Limestone Formation	Ireland	A member of Orthida belonging to the family Platystrophiidae.
Pustulobolus^[50]	Gen. et sp. nov	Valid	Skovsted et al.	Cambrian Stage 3-4	Forteau Formation	Canada ( Newfoundland and Labrador)	A member of Linguloidea belonging to the family Eoobolidae. The type species is P. triangulus.
Qidongia^[59]	Gen. et sp. nov	Valid	Lü & Ma	Devonian (late Frasnian)		China	A member of Terebratulida. The type species is Q. tani.
Rhipidomella discreta^[76]	Sp. nov	Valid	Cisterna et al.	Carboniferous (late Serpukhovian–Bashkirian)	El Paso Formation	Argentina	A brachiopod belonging to the group Orthida and the family Rhipidomellidae.
Rioultina zalasensis^[77]	Sp. nov	Valid	Radwańska	Late Jurassic (Oxfordian)		Poland	A member of Thecideida belonging to the family Thecidellinidae.
Sericoidea hibernica^[75]	Sp. nov	Valid	Harper, Parkes & Zhan	Ordovician (Katian)	Raheen Formation	Ireland	A plectambonitoid brachiopod belonging to the family Sowerbyellidae.
Serratocrista scaldisensis^[71]	Sp. nov	Valid	Mottequin & Simon	Carboniferous (Tournaisian)	Tournai Formation	Belgium	A member of Orthotetida belonging to the family Schuchertellidae.
Simehorthis^[78]	Gen. et sp. nov	Valid	Kebria-Ee Zadeh, Popov & Ghobadi Pour	Ordovician (Darriwilian)	Lashkarak Formation	Iran	A member of Orthida belonging to the family Hesperorthidae. Genus includes new species S. fascicostellata.
Somalithyris lakhaparensis^[79]	Sp. nov	Valid	Mukherjee & Shome	Late Jurassic (Tithonian)		India
Starnikoviella^[54]	Gen. et sp. nov	Valid	Baranov	Ordovician		Russia	The type species is S. settedabanica.
Tectogonotoechia rivasi^[80]	Sp. nov	Valid	García-Alcalde & Herrera	Devonian (Pragian)	Nogueras Formation	Spain	A member of Rhynchonellida belonging to the superfamily Ancistrorhynchoidea and the family Iberirhynchiidae.
Thomasaria? baii^[59]	Sp. nov	Valid	Lü & Ma	Devonian (late Frasnian)		China	A member of Spiriferida.
Thomasaria? liangi^[59]	Sp. nov	Valid	Lü & Ma	Devonian (late Frasnian)		China	A member of Spiriferida.
Tunethyris blodgetti^[81]	Sp. nov	Valid	Feldman	Middle Triassic	Saharonim Formation	Israel	A member of Terebratulida belonging to the family Dielasmatidae.
Westonia mardini^[82]	Sp. nov	Valid	Mergl et al.	Cambrian (Furongian)	Sosink Formation	Turkey
Xiangia^[59]	Gen. et sp. nov	Junior homonym	Lü & Ma	Devonian (late Frasnian)		China	A member of Spiriferida. The type species is X. liaoi. The generic name is preoccupied by Xiangia Peng (1987).
Zhanorthis^[62]	Gen. et sp. nov	Valid	Popov & Cocks	Ordovician (Dapingian)		Iran	An orthoid brachiopod. Genus includes new species Z. gerdkuhensis.
Ziyunospirifer^[83]	Nom. nov	Valid	Shen in Shen et al.	Early Carboniferous	Zhaojiashan Formation	China	A member of Spiriferida belonging to the family Choristitidae; a replacement name for Quizhouspirifer Xian (1982). The type species is "Quizhouspirifer" ziyunensis Xian (1982).

Molluscs

Main article: 2017 in paleomalacology

Echinoderms

Research

Systematic revision of the North American members of the diploporitan family Holocystitidae is published by Sheffield & Sumrall (2017).^[84]
Triassic members of the otherwise Paleozoic groups of sea urchins (the family Proterocidaridae), brittle stars (the family Eospondylidae) and starfish are reported by Thuy, Hagdorn & Gale (2017).^[85]^[86]^[87]^[88]^[89]^[90]^[91]
Phylogenetic analysis and systematic revision of early to middle Paleozoic non-camerate crinoids published by Wright (2017).^[92]
Systematic revision of Ordovician camerate crinoids published by Cole (2017).^[93]
Major revision to the classification of fossil and extant Crinoidea by Wright et al. (2017), including the presentation of new phylogeny-based and rank-based classifications.^[94]
A study on large-scale patterns of morphologic evolution in the Paleozoic radiation of eucladid crinoids is published by Wright (2017).^[95]
A study on the internal morphology of the water vascular system in a specimen of a stem-ophiuroid species Protasterina flexuosa from the Ordovician (Katian) Kope Formation (Kentucky, United States) is published by Clark et al. (2017).^[96]
A study on the paleoecology of the echinoderm species known from the upper Campanian Pierre Shale (including the crinoid Lakotacrinus brezinai), especially on their adaptations to the cold seep environment, is published by Kato, Oji & Shirai (2017).^[97]^[98]^[99]

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Amblypygus matruhensis^[100]	Sp. nov	Valid	Ali	Middle Miocene		Egypt	A sea urchin.
Ambonacrinus^[101]	Gen. et sp. nov	Valid	Cole et al.	Ordovician (Katian)	Fombuena Formation	Spain	A diplobathrid camerate crinoid. Genus includes new species A. decorus.
Andymetra toarcensis^[102]	Sp. nov	Valid	Hess & Thuy	Early Jurassic		France	A comatulid crinoid.
Anthroosasterias^[103]	Gen. et sp. nov	Valid	Blake	Carboniferous	Gilmore City Formation	United States ( Iowa)	A starfish belonging to the family Urasterellidae. Genus includes new species A. mikrotero.
Antillaster farisi^[104]	Sp. nov	Valid	Ali	Middle Eocene		Egypt	A sea urchin.
Aspidophiura? seren^[105]	Sp. nov	Valid	Ewin & Thuy	Jurassic	Oxford Clay Formation	United Kingdom	A brittle star.
Ateleocystites? lansae^[106]	Sp. nov	Valid	McDermott & Paul	Ordovician (Katian)	Slade and Redhill Beds	United Kingdom	A mitrate belonging to the family Anomalocystitidae, possibly a species of Ateleocystites.
Brissus mihalyi^[107]	Sp. nov.	Valid	Polonkai et al.	Middle Miocene	Leitha Limestone Formation	Hungary	A heart urchin belonging to the family Brissidae.
Crepidosoma doylei^[108]	Sp. nov	Valid	Blake, Donovan & Harper	Silurian (Telychian)	Kilbride Formation	Ireland	A brittle star belonging to the group Oegophiurida and the family Encrinasteridae.
Dalicrinus^[101]	Gen. et sp. nov	Valid	Cole et al.	Ordovician (Katian)	Fombuena Formation	Spain	A diplobathrid camerate crinoid. Genus includes new species D. hammanni.
Diplodetus brisenoi^[109]	Sp. nov	Valid	Silva-Martínez et al.	Late Cretaceous (early Campanian)	Austin Formation	Mexico	A heart urchin belonging to the family Brissidae.
Echinocyamus belali^[104]	Sp. nov	Valid	Ali	Middle Eocene		Egypt	A sea urchin.
Enakomusium whymanae^[105]	Sp. nov	Valid	Ewin & Thuy	Jurassic	Oxford Clay Formation	United Kingdom	A brittle star.
Eopatelliocrinus hispaniensis^[101]	Sp. nov	Valid	Cole et al.	Ordovician (Katian)	Fombuena Formation	Spain	A monobathrid camerate crinoid.
Eotiaris guadalupensis^[110]	Sp. nov	Valid	Thompson in Thompson, Petsios & Bottjer	Permian (Capitanian)	Bell Canyon Formation	United States ( Texas)	A sea urchin. The name first appeared in the publication of Thompson et al. (2015);^[111] however, it was published in an online only journal Scientific Reports and it was not registered with ZooBank, making it invalid until it was validated by Thompson, Petsios & Bottjer (2017).^[110]
Felbabkacystis^[112]	Gen. et sp. nov	Valid	Nardin et al.	Cambrian (Drumian)	Jince Formation	Czech Republic	A transitional form between calyx-bearing and theca-bearing blastozoans. Genus includes new species F. luckae.
Fombuenacrinus^[101]	Gen. et sp. nov	Valid	Cole et al.	Ordovician (Katian)	Fombuena Formation	Spain	A diplobathrid camerate crinoid. Genus includes new species F. nodulus.
Forcipicrinus^[102]	Gen. et sp. nov	Valid	Hess & Thuy	Early Jurassic		France	An isocrinid crinoid. Genus includes new species F. normannicus.
Globator roselli^[113]	Sp. nov	Valid	Carrasco	Eocene		Spain	A sea urchin related to members of the genus Conulus.
Goniopygus emmae^[114]	Sp. nov	Valid	Forner i Valls	Late Cretaceous (Campanian)		Morocco	A sea urchin belonging to the group Arbacioida and the family Acropeltidae.
Grigopyrgus^[115]	Gen. et comb. nov	Valid	Müller & Hahn	Early Devonian		Germany	A member of Edrioasteroidea belonging to the family Agelacrinitidae; a new genus for "Agelacrinites" curvatus Grigo (1995).
Goyacrinus^[101]	Gen. et sp. nov	Valid	Cole et al.	Ordovician (Katian)	Fombuena Formation	Spain	A diplobathrid camerate crinoid. Genus includes new species G. gutierrezi.
Heropyrgus^[116]	Gen. et sp. nov	Valid	Briggs et al.	Silurian	Herefordshire Lagerstätte	United Kingdom	A rhenopyrgid edrioasteroid. The type species is H. disterminus.
Holocystites salmoensis^[117]	Sp. nov	Valid	Sheffield, Ausich & Sumrall	Ordovician (Hirnantian)	Ellis Bay Formation	Canada ( Quebec)	A member of Diploporita belonging to the group Sphaeronitida and the family Holocystitidae.
Metalia lindaae^[104]	Sp. nov	Valid	Ali	Middle Eocene		Egypt	A sea urchin.
Monostychia alanrixi^[118]	Sp. nov	Valid	Sadler, Martin & Gallagher	Miocene	Colville Sandstone	Australia	A sea urchin.
Monostychia macnamarai^[118]	Sp. nov	Valid	Sadler, Martin & Gallagher	Miocene	Colville Sandstone	Australia	A sea urchin.
Monostychia robertirwini^[118]	Sp. nov	Valid	Sadler, Martin & Gallagher	Miocene	Colville Sandstone	Australia	A sea urchin.
Moroccodiscus^[119]	Gen. et sp. nov	Valid	Reich et al.	Ordovician (Darriwilian)	Taddrist Formation	Morocco	A cyclocystoid echinoderm. Genus includes new species M. smithi.
Oehlerticrinus peachi^[120]	Sp. nov	Valid	Donovan & Fearnhead	Early Devonian	Looe Basin	United Kingdom	A crinoid belonging to the group Monobathrida and the family Hexacrinitidae.
Ophiotitanos smithi^[105]	Sp. nov	Valid	Ewin & Thuy	Jurassic	Oxford Clay Formation	United Kingdom	A brittle star.
Ova rancoca^[121]	Sp. nov	Valid	Zachos	Paleocene (Thanetian)	Vincentown Formation	United States ( New Jersey)	A sea urchin.
Paerticrinus^[122]	Gen. et sp. nov	Valid	Wright & Toom	Silurian (Rhuddanian)		Estonia	A crinoid. Genus includes new species P. arvosus.
Palaeocomaster structus^[102]	Sp. nov	Valid	Hess & Thuy	Early Jurassic		France	A comatulid crinoid.
Persiacarpos^[123]	Gen. et sp. nov	Valid	Rozhnov & Parsley	Cambrian	Mila Formation	Iran	A member of Cornuta. Genus includes new species P. jefferiesi.
Petalobrissus ossoi^[114]	Sp. nov	Valid	Forner i Valls	Late Cretaceous (Campanian)		Morocco	A sea urchin belonging to the group Cassiduloida and the family Faujasidae.
Petalocrinus stenopetalus^[124]	Sp. nov	Valid	Mao et al.	Silurian (Aeronian)		China	A crinoid belonging to the family Petalocrinidae.
Picassocrinus^[101]	Gen. et sp. nov	Valid	Cole et al.	Ordovician (Katian)	Fombuena Formation	Spain	A cladid crinoid. Genus includes new species P. villasi.
Ronsocrinus^[125]	Gen. et sp. nov	Valid	Cordie & Witzke	Devonian (Givetian)		United States ( Iowa)	A camerate crinoid belonging to the family Melocrinitidae. Genus includes new species R. rabia.
Salenia palmyra^[121]	Sp. nov	Valid	Zachos	Paleocene (Danian)	Clayton Formation	United States ( Alabama Georgia (U.S. state))	A sea urchin.
Sanducystis^[126]	Gen. et sp. nov	Valid	Zamora et al.	Cambrian (Furongian)	Sandu Formation	China	A stemmed echinoderm. The type species is S. sinensis.
Singillatimetra truncata^[102]	Sp. nov	Valid	Hess & Thuy	Early Jurassic		France	An isocrinid crinoid.
Solanocrinites jagti^[102]	Sp. nov	Valid	Hess & Thuy	Early Jurassic		France	A comatulid crinoid.
Spinimetra^[102]	Gen. et sp. nov	Valid	Hess & Thuy	Early Jurassic		France	A comatulid crinoid. Genus includes new species S. chesnieri.
Spirocrinus circularis^[124]	Sp. nov	Valid	Mao et al.	Silurian (Aeronian)		China	A crinoid belonging to the family Petalocrinidae.
Spirocrinus dextrosus^[124]	Sp. nov	Valid	Mao et al.	Silurian (Aeronian)		China	A crinoid belonging to the family Petalocrinidae.
Staurasterias^[103]	Gen. et sp. nov	Valid	Blake	Carboniferous	Keokuk Formation	United States ( Indiana)	A starfish belonging to the family Urasterellidae. Genus includes new species S. elegans.
Sumrallia^[127]	Gen. et sp. nov	Valid	Müller & Hahn	Early Devonian	Seifen Formation	Germany	A member of Edrioasteroidea. Genus includes new species S. rseiberti.
Superstesaster^[128]	Gen. et sp. nov	Valid	Villier et al.	Early Triassic		United States ( Utah)	A starfish. Genus includes new species S. promissor.
Teleosaster^[129]	Gen. et sp. nov	Valid	Hunter & McNamara	Permian (Kungurian)	Cundlego Formation	Australia	A brittle star. Genus includes new species T. creasyi.
Tintinnabulicrinus^[122]	Gen. et sp. nov	Valid	Wright & Toom	Ordovician (Katian)		Estonia	A crinoid. Genus includes new species T. estoniensis.
Ulphaceaster^[130]	Gen. et sp. nov	Valid	Néraudeau et al.	Late Cretaceous (Cenomanian)		France	A sea urchin belonging to the family Archiaciidae. Genus includes new species U. sarthacensis.
Vologesia rollingstones^[131]	Sp. nov	Valid	Schlüter & Wiese	Late Cretaceous (early Campanian)		Spain	A sea urchin belonging to the family Echinolampadidae.

Conodonts

Research

A study on the conodont assemblage from the Silurian (Homerian) Rootsiküla Formation (Estonia), interpreted as occurring in the evaporite-bearing strata, and on the conodont diversity in various environments, is published by Jarochowska et al. (2017).^[132]
Articulated skeletal remains of Hindeodus parvus, providing direct evidence of the number and arrangement of elements in the apparatus, are described from the Lower Triassic of China by Zhang et al. (2017).^[133]^[134]^[135]

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Acodus zeballus^[136]	Sp. nov	Valid	Voldman & Albanesi in Voldman et al.	Early Ordovician		Argentina
Aldridgeognathus^[137]	Gen. et sp. nov	Valid	Miller et al.	Ordovician (Darriwilian)	Amdeh Formation	Oman	A member of Balognathidae. Genus includes new species A. manniki.
Bispathodus ultimus corradinii^[138]	Subsp. nov	Valid	Söte, Hartenfels & Becker	Devonian (Famennian)		Germany
Coelocerodontus hunanensis^[139]	Sp. nov	Valid	Dong & Zhang	Cambrian (Furongian)	Panjiazui Formation	China	A euconodont.
Ctenopolygnathus parallelus^[140]	Sp. nov	Valid	Ovnatanova et al.	Late Devonian	Kedzyrschor Formation	Russia
Fahraeusodus jachalensis^[141]	Sp. nov	Valid	Feltes & Albanesi in Serra et al.	Ordovician (Darriwilian)	Gualcamayo Formation Las Aguaditas Formation Las Chacritas Formation San Juan Formation	Argentina
Furnishina wangcunensis^[139]	Sp. nov	Valid	Dong & Zhang	Cambrian (Furongian)	Bitiao Formation	China	A member of Paraconodontida.
Gothodus vetus^[136]	Sp. nov	Valid	Voldman & Albanesi in Voldman et al.	Early Ordovician		Argentina
Guexispathodus^[142]	Gen. et comb. nov	Valid	Plasencia et al.	Middle Triassic	Mukheiris Formation Saharonim Formation	Israel Jordan	A member of the family Gondolellidae. The type species is "Neospathodus" shagami Benjamini & Chepstow-Lusty (1986); genus also includes "Pseudofurnishius" siyalaensis Sadeddin & Kozur (1992).
Gullodus tieqiaoensis^[143]	Sp. nov	Valid	Sun et al.	Permian		China
Icriodus ballbergensis^[144]	Sp. nov	Valid	Lüddecke, Hartenfels & Becker	Devonian (Famennian)		Germany
Icriodus marieae^[145]	Sp. nov	Valid	Suttner, Kido & Suttner	Middle Devonian	Valentin Formation	Austria France Germany
Idiognathodus boardmani^[146]	Sp. nov	Valid	Hogancamp & Barrick	Carboniferous (Gzhelian)	Heebner Shale	United States
Idiognathodus itaitubensis^[147]	Sp. nov	Valid	Cardoso, Sanz-López & Blanco-Ferrera	Carboniferous (Pennsylvanian)	Tapajós Group	Brazil
Idiognathoides luokunensis^[148]	Sp. nov	Valid	Hu & Qi in Hu et al.	Carboniferous (Bashkirian)		China
Iowagnathus^[149]	Gen. et sp. nov	Valid	Liu et al.	Ordovician (Whiterock Stage)	Winneshiek Konservat-Lagerstätte	United States ( Iowa)	Genus includes new species I. grandis.
Kirilella^[142]	Gen. et comb. nov	Valid	Plasencia et al.	Middle Triassic		Austria Canada China Egypt Hungary Israel Italy Japan Jordan Russia Spain United States	A member of the family Gondolellidae. The type species is "Polygnathus" mungoensis Diebel (1956); genus also includes "Tardogondolella" diebeli Kozur & Mostler (1971), "Epigondolella" mostleri Kozur in Kozur & Mock (1972) and "Metapolygnathus" longobardicus Kovács (1983).
Laiwugnathus hunanensis^[139]	Sp. nov	Valid	Dong & Zhang	Cambrian (Drumian)	Huaqiao Formation	China	A member of Paraconodontida.
Laiwugnathus transitans^[139]	Sp. nov	Valid	Dong & Zhang	Cambrian (Guzhangian and Paibian)	Chefu Formation	China	A member of Paraconodontida.
Lenathodus^[150]	Gen. et sp. nov	Valid	Izokh in Izokh & Yazikov	Early Carboniferous		Russia	Genus includes new species L. bakharevi.
Lugnathus^[139]	Gen. et sp. nov	Valid	Dong & Zhang	Cambrian Stage 10 and Early Ordovician (Tremadocian)	Panjiazui Formation	China	A member of Paraconodontida. Genus includes new species L. hunanensis.
Marquezella^[142]	Gen. et comb. nov	Valid	Plasencia et al.	Middle Triassic		Austria Bulgaria China France Greece Hungary India Italy Japan Russia Slovakia Slovenia Spain	A member of the family Gondolellidae. The type species is "Gladigondolella" truempyi Hirsch (1971); genus also includes "Polygnathus" japonicus Hayashi (1968).
Mayrodus^[151]	Gen. et sp. nov	Valid	Zhang, Jowett & Barnes	Silurian (Sheinwoodian)	Cape Phillips Formation	Canada ( Nunavut)	A conodont of uncertain phylogenetic placement. The type species is M. melchini.
Miaognathus^[139]	Gen. et sp. nov	Valid	Dong & Zhang	Cambrian Stage 10	Shenjiawan Formation	China	A member of Paraconodontida. Genus includes new species M. multicostatus.
Millerodontus^[139]	Gen. et sp. nov	Valid	Dong & Zhang	Cambrian (Furongian)	Shenjiawan Formation	China	A euconodont. Genus includes new species M. intermedius.
Mosherella praebudaensis^[152]	Sp. nov	Valid	Chen & Lukeneder	Late Triassic (Carnian)	Kasimlar Formation	Turkey
Neopolygnathus communis yazikovi^[150]	Subsp. nov	Valid	Izokh in Izokh & Yazikov	Early Carboniferous		Russia
Neopolygnathus crucesignatis^[153]	Sp. nov	Valid	Plotitsyn & Zhuravlev	Carboniferous (Tournaisian)		Russia
Norigondolella carlae^[154]	Sp. nov	In press	Rigo et al.	Late Triassic (Carnian)	Scillato Formation	Austria Italy Turkey	A member of Ozarkodinida.
Omanognathus^[137]	Gen. et sp. nov	Valid	Miller et al.	Ordovician (Darriwilian)	Amdeh Formation	Oman	A member of Balognathidae. Genus includes new species O. daiqaensis.
Palmatolepis chernovi^[155]	Sp. nov	Valid	Soboleva	Devonian (Frasnian)		Russia
Palmatolepis spallettae^[156]	Nom. nov	Valid	Klapper et al.	Devonian (Frasnian)		Canada ( Ontario)	A replacement name for Palmatolepis nodosa Klapper et al. (2004).
Palmatolepis zhuravlevi^[155]	Sp. nov	Valid	Soboleva	Devonian (Frasnian)		Russia
Polygnathus arcus^[153]	Sp. nov	Valid	Plotitsyn & Zhuravlev	Carboniferous (Tournaisian)		Russia
Polygnathus mawsonae^[140]	Sp. nov	Junior homonym	Ovnatanova et al.	Devonian (Famennian)	Sortomael' Formation	Australia Russia	Ovnatanova et al. (2019) coined a replacement name Polygnathus sharyuensis.^[157]
Polygnathus postvogesi^[158]	Sp. nov	Valid	Plotitsyn & Zhuravlev	Carboniferous (Tournaisian)		Russia
Prosagittodontus compressus^[139]	Sp. nov	Valid	Dong & Zhang	Cambrian (Guzhangian and Paibian)	Chefu Formation	China	A member of Paraconodontida.
Pseudohindeodus elliptica^[143]	Sp. nov	Valid	Sun et al.	Permian		China
Quadralella wanlanensis^[159]	Sp. nov	Valid	Zhang et al.	Triassic		China
Quadralella yongyueensis^[159]	Sp. nov	Valid	Zhang et al.	Triassic		China
Siphonodella carinata^[160]	Sp. nov	Valid	Zhuravlev	Carboniferous (Tournaisian)	Idzhid Formation	Russia ( Komi Republic)
Siphonodella kalvodai^[161]	Sp. nov	Valid	Kaiser, Kumpan & Cígler	Carboniferous (Tournaisian)	Líšeň Formation	Czech Republic Tajikistan	A member of Ozarkodinida belonging to the family Elictognathidae.
Sweetognathus asymmetrica^[143]	Sp. nov	Valid	Sun et al.	Permian		China
Tujiagnathus^[139]	Gen. et sp. nov	Valid	Dong & Zhang	Cambrian (Furongian)	Bitiao Formation	China	A euconodont. Genus includes new species T. gracilis.
Vjalovognathus carinatus^[162]	Sp. nov	Valid	Wang et al.	Permian (Changhsingian)		China India
Wangcunella^[139]	Gen. et sp. nov	Valid	Dong & Zhang	Cambrian (Furongian)	Bitiao Formation	China	A euconodont. Genus includes new species W. conicus.
Wangcunognathus^[139]	Gen. et sp. nov	Valid	Dong & Zhang	Cambrian (Paibian)	Bitiao Formation	China	A member of Paraconodontida. Genus includes new species W. elegans.
Westergaardodina dimorpha^[139]	Sp. nov	Valid	Dong & Zhang	Cambrian (Paibian)	Bitiao Formation	China	A member of Paraconodontida.
Westergaardodina gigantea^[139]	Sp. nov	Valid	Dong & Zhang	Cambrian (Guzhangian)	Chefu Formation	China	A member of Paraconodontida.
Westergaardodina sola^[139]	Sp. nov	Valid	Dong & Zhang	Cambrian (Guzhangian)	Chefu Formation	China	A member of Paraconodontida.
Zentagnathus^[136]	Gen. et comb. nov	Valid	Voldman & Albanesi in Voldman et al.	Early Ordovician		Argentina	A new genus for "Trapezognathus" primitivus Voldman, Albanesi & Zeballo in Voldman et al. (2013); genus also includes "Trapezognathus" argentinensis Rao et al. (1994)

Fishes

Main article: 2017 in paleoichthyology

Amphibians

Research

A study on the evolution of eye size in early tetrapods and in fish belonging to the lineage that gave rise to tetrapods, as well as on the impact of the eye size on the eye performance while viewing objects through water and through air is published by MacIver et al. (2017).^[163]
A study on the evolution of forelimb musculature from the lobe-finned fish to early tetrapods is published online by Molnar et al. (2017).^[164]
A study on the influence of habitat traits on the persistence length of living and fossil amphibian species is published by Tietje & Rödel (2017).^[165]
A study on the development of the vertebral intercentrum and pleurocentrum in fossil amphibians is published by Danto et al. (2017).^[166]
A study on the probable function of the interpterygoid vacuities (holes in the palate) in temnospondyls as the site of muscle attachment is published by Witzmann & Werneburg (2017).^[167]
A study on the earliest larval development in temnospondyls, as indicated by specimens from the Permian (Sakmarian) lake sediments near Obermoschel (Saar–Nahe Basin, Germany), is published by Werneburg (2017).^[168]
A study on the histology of the small palatal plates and their denticles in a Permian dissorophoid temnospondyl from the Dolese Brothers Limestone Quarry near Richards Spur (Oklahoma, United States) is published by Gee, Haridy & Reisz (2017).^[169]
Taxonomic revision of all described rhinesuchids and a study on the phylogenetic relationships of members of Rhinesuchidae is published by Marsicano et al. (2017), who transfer the species "Rhinesuchus" capensis Haughton (1925) to the genus Rhinesuchoides.^[170]
New specimen of the rhinesuchid Australerpeton cosgriffi (a skull and mandible) is described from the Permian Rio do Rasto Formation (Brazil) by Azevedo, Vega & Soares (2017).^[171]
A description of the anatomy of the braincase and middle ear regions of an exceptionally well-preserved skull of Stanocephalosaurus amenasensis from the Triassic of Algeria is published by Arbez, Dahoumane & Steyer (2017).^[172]
A study on the anatomy of the skulls of metoposaurid species Metoposaurus krasiejowensis and Apachesaurus gregorii, as well as its implications for establishing whether metoposaurids were active or ambush predators is published by Fortuny, Marcé-Nogué & Konietzko-Meier (2017).^[173]
An analysis of the microanatomy and histology of metoposaurid vertebra from the Petrified Forest National Park is published by Gee, Parker & Marsh (2017), who interpret Apachesaurus gregorii as more likely to be an early ontogenetic stage of a large metoposaurid, such as Koskinonodon perfectus rather than a distinct species.^[174]
A juvenile specimen of Koskinonodon perfectus is described from the Norian Petrified Forest Member of the Late Triassic Chinle Formation (Arizona, United States) by Gee & Parker (2017).^[175]
A study on the physiology (especially metabolic rate, body temperature, breathing, feeding, digestion, osmoregulation and excretion) of Archegosaurus decheni is published by Witzmann & Brainerd (2017).^[176]
A study on the histology of the dermal skull roof bones in Kokartus honorarius is published by Skutschas & Boitsova (2017).^[177]
Fossilized soft tissues preserved with the type specimen of the salamander Phosphotriton sigei are described by Tissier, Rage & Laurin (2017).^[178]
A study on the bite force in extant Cranwell's horned frog (Ceratophrys cranwelli) and its implications for estimating the bite force in the Late Cretaceous species Beelzebufo ampinga is published by Lappin et al. (2017).^[179]
Frog fossils, including the first known fossils of shovelnose frogs, are described from the early Pliocene of Kanapoi (Kenya) by Delfino (2017).^[180]
A study on the morphology of the skull of Lethiscus stocki and on the phylogenetic relationships of early tetrapods, recovering lepospondyls as a polyphyletic group, is published by Pardo et al. (2017).^[181]

New taxa

Temnospondyls

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Aphaneramma gavialimimus^[182]	Sp. nov	Valid	Fortuny et al.	Early Triassic (Olenekian)		Madagascar		Aphaneramma
Chinlestegophis^[183]	Gen. et sp. nov	Valid	Pardo, Small & Huttenlocker	Late Triassic	Chinle Formation	United States ( Colorado)	A member of Stereospondyli, possibly a stem-caecilian. The type species is C. jenkinsi.
Cyclotosaurus naraserluki^[184]	Sp. nov	Valid	Marzola et al.	Late Triassic	Fleming Fjord Formation	Greenland		Cyclotosaurus
Tomeia^[185]	Gen. et sp. nov	Valid	Eltink, Stock Da-Rosa, & Dias-da-Silva	Early Triassic	Sanga do Cabral Formation	Brazil	A capitosaur.

Lissamphibians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Chachaiphrynus^[186]	Gen. et sp. nov	Valid	Nicoli	Oligocene		Argentina	A member of Odontophrynidae. The type species is C. lynchi.
Genibatrachus^[187]	Gen. et sp. nov	Valid	Gao & Chen	Early Cretaceous	Guanghua (upper part of Longjiang) Formation	China	A crown-group frog. The type species is G. baoshanensis.
Sanshuibatrachus^[188]	Gen. et sp. nov	Valid	Wang, Roček & Dong	Early Eocene		China	A pelobatoid frog of uncertain phylogenetic placement. Genus includes new species S. sinensis.

Other amphibians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Spathicephalus marsdeni^[189]	Sp. nov	Valid	Smithson et al.	Carboniferous (Viséan)	Anstruther Formation	United Kingdom	A member of the superfamily Baphetoidea.
Yumenerpeton^[190]	Gen. et sp. nov	Valid	Jiang, Ji & Mo	Middle Permian	Xidagou Formation	China	A bystrowianid chroniosuchian. The type species is Y. yangi.

Reptiles

Main articles: 2017 in reptile paleontology and 2017 in archosaur paleontology

Synapsids

Non-mammalian synapsids

Research

Phreatophasma aenigmaticum is argued to be a member of Caseidae by Brocklehurst & Fröbisch (2017).^[191]
New fossil material of the caseid Alierasaurus ronchii is described from the Permian deposits of Cala del Vino Formation (Sardinia, Italy) by Romano et al. (2017).^[192]
A study on the histology of the humeri of Ophiacodon, revealing the existence of fibrolamellar bone in the postcranial bones of this taxon, is published by Shelton & Sander (2017).^[193]
A study on the body size evolution of edaphosaurids and sphenacodontids is published by Brocklehurst & Brink (2017).^[194]
A study on the evolution of the endothermy in non-mammalian therapsids as indicated by oxygen isotope composition of bone and tooth phosphate in Permian and Triassic therapsids is published by Rey et al. (2017).^[195]
A study on the brain morphology of non-mammaliaform therapsids based on skull endocasts of Moschops capensis and a number of biarmosuchians (including Herpetoskylax hopsoni and members of the genera Hipposaurus and Lemurosaurus) is published by Benoit et al. (2017).^[196]
A study on the morphology of the bony labyrinth of five biarmosuchian specimens is published by Benoit et al. (2017).^[197]
A study on the anatomy of the skull of Moschops capensis, revealing adaptations of the central nervous system related to head-to-head fighting, is published by Benoit et al. (2017).^[198]
A study on the resting metabolic rate in Moghreberia nmachouensis is published by Olivier et al. (2017).^[199]
A study on the contents of the depression known as the "unossified zone" in the brain cavity of Diictodon feliceps is published by Laaß, Schillinger & Kaestner (2017).^[200]
A reassessment of the skull morphology and phylogenetic position of Compsodon helmoedi is published by Angielczyk & Kammerer (2017).^[201]
A skeleton of Lystrosaurus curvatus in a fossilized burrow, preserved with taphonomic evidence indicating that this individual was the burrow maker, is described from the Lower Triassic of the South African Karoo Basin by Botha-Brink (2017).^[202]
A structure analogous to the mammalian neocortex is reported in Kawingasaurus fossilis by Laaß & Kaestner (2017).^[203]
A gorgonopsian dentary affected by a condition closely resembling compound odontoma is reported from the Upper Permian of Tanzania by Whitney, Mose & Sidor (2017).^[204]
A detailed description of the braincase of two gorgonopsian specimens (a probable specimen of Aelurosaurus wilmanae from South Africa and a possible specimen of Arctognathus? nasuta from Tanzania) is published by Araújo et al. (2017).^[205]
A redescription and revision of the gorgonopsian genus Arctops is published by Kammerer (2017).^[206]
Rediscovered holotype of the gorgonopsian species Clelandina major is described by Kammerer (2017), who considers this species to be a junior synonym of Clelandina rubidgei.^[207]
A study on the anatomy of the teeth and maxilla of Euchambersia mirabilis and its implications for the hypothesis that venom gland were present in this species is published by Benoit et al. (2017).^[208]
A redescription and a study on the phylogenetic relationships of Silphoictidoides ruhuhuensis is published by Maisch (2017), who considers the species to be a basal member of Baurioidea.^[209]
A study on the internal morphology of the interorbital region of the skull of basal cynodonts, including rarely fossilized orbitosphenoid elements, is published by Benoit et al. (2017).^[210]
A study on the anatomy of the nasal regions of the non-mammalian cynodonts Massetognathus, Probainognathus and Elliotherium, comparing it to the nasal regions of fossil mammaliaforms and extant mammals, is published by Crompton et al. (2017).^[211]
A survey of the aggregations of the specimens of Galesaurus planiceps and Thrinaxodon liorhinus, with emphasis on whether the aggregations consist of individuals of similar age or representing a mixture of different age classes, is published by Jasinoski & Abdala (2017).^[212]
A study on the ontogenetic changes in the skull and mandible of Galesaurus planiceps is published by Jasinoski & Abdala (2017).^[213]
A description of the postcranial skeleton of Boreogomphodon from the Triassic Pekin Formation (North Carolina, United States) and a review of the postcranial variation across members of the family Traversodontidae is published by Liu, Schneider & Olsen (2017).^[214]
A study on the jaw movement of Exaeretodon argentinus as indicated by its dental microwear is published by Kubo, Yamada & Kubo (2017).^[215]
A study on the morphology of the teeth of the cynodont Candelariodon barberenai, as well as on the phylogenetic relationships of the species, is published by Martinelli et al. (2017).^[216]
A description of the anatomy of the postcranial skeleton of Tritylodon longaevus is published by Gaetano, Abdala & Govender (2017).^[217]
A reassessment of the anatomy of the postcanine teeth of Stereognathus, based upon all available material from the United Kingdom, is published by Panciroli et al. (2017), who consider the species S. hebridicus to be a junior synonym of the species S. ooliticus.^[218]
Cast of a burrow which was probably made by a tritheledontid cynodont is described from the Early Jurassic upper Elliot Formation (South Africa) by Bordy et al. (2017).^[219]
A study on the evolution of jaw muscles across the cynodont–mammaliaform transition is published by Lautenschlager et al. (2017).^[220]

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Alemoatherium^[221]	Gen. et sp. nov	Valid	Martinelli et al.	Late Triassic (late Carnian)	Santa Maria Formation	Brazil	A cynodont belonging to the group Prozostrodontia. The type species is A. huebneri.
Aleodon cromptoni^[222]	Sp. nov	Valid	Martinelli et al.	Triassic (Ladinian—early Carnian)		Brazil Namibia?	A cynodont belonging to the family Chiniquodontidae.
Bulbasaurus^[223]	Gen. et sp. nov	Valid	Kammerer & Smith	Late Permian	Teekloof Formation	South Africa	A dicynodont belonging to the family Geikiidae. The type species is B. phylloxyron.
Dalongkoua^[224]	Gen. et sp. nov	Valid	Liu & Abdala	Late Permian	Guodikeng Formation	China	A therocephalian. The type species is D. fuae.
Microwhaitsia^[225]	Gen. et sp. nov	Valid	Huttenlocker & Smith	Permian (Wuchiapingian)	Teekloof Formation	South Africa	A whaitsiid therocephalian. The type species is M. mendrezi.
Nuurtherium^[226]	Gen. et sp. nov	Valid	Velazco, Buczek & Novacek	Late Jurassic	Ulan Malgait Sequence	Mongolia	A tritylodontid cynodont. The type species is N. baruunensis.
Ophidostoma^[225]	Gen. et sp. nov	Valid	Huttenlocker & Smith	Permian (Wuchiapingian)	Teekloof Formation	South Africa	A whaitsioid therocephalian of uncertain phylogenetic placement. The type species is O. tatarinovi.
Parasuminia^[227]	Gen. et sp. nov	Valid	Kurkin	Permian (Severodvinian)	Poldarsa Formation	Russia	An anomodont related to Suminia. Genus includes new species P. ivakhnenkoi.
Scalenodon ribeiroae^[228]	Sp. nov	Valid	Melo, Martinelli & Soares	Triassic	Santa Maria Supersequence	Brazil	A traversodontid cynodont.
Shartegodon^[226]	Gen. et sp. nov	Valid	Velazco, Buczek & Novacek	Late Jurassic	Ulan Malgait Sequence	Mongolia	A tritylodontid cynodont. The type species is S. altai.
Shiguaignathus^[229]	Gen. et sp. nov	Valid	Liu & Abdala	Late Permian	Naobaogou Formation	China	An akidnognathid therocephalian. The type species is S. wangi.

Mammals

Main article: 2017 in mammal paleontology

Other animals

Research

A study on a succession of Ediacaran to Cambrian fossil assemblages from the eastern Siberian Platform (Russia) is published by Zhu et al. (2017), who argue that so-called Ediacaran and earliest Cambrian skeletal biotas overlap without notable biotic turnover.^[230]
A study on the Ediacaran taxon Parvancorina minchami, indicating that this animal was capable of performing rheotaxis, is published by Paterson et al. (2017).^[231]
A study on the water flow around the body of the Ediacaran taxon Parvancorina and its implications for the feeding mode and mobility of this animal is published by Darroch et al. (2017).^[232]
Fossils of members of the genus Namacalathus (co-occurring with Cloudina and Corumbella) are reported from the Ediacaran Tagatiya Guazú Formation (Itapucumi Group, Paraguay) by Warren et al. (2017), extending known geographic range of the taxon.^[233]
A study on the morphology, growth and development of Dickinsonia costata is published by Evans, Droser & Gehling (2017).^[234]
A study on the growth and development of Dickinsonia is published by Hoekzema et al. (2017), who interpret this taxon as an animal.^[235]
A study on the anatomy of Dickinsonia costata and D. tenuis is published by Zakrevskaya & Ivantsov (2017), who interpret D. costata as probably descended from D. tenuis by neoteny.^[236]
Description of newly discovered disc-shaped, soft-bodied fossils from the early Cambrian Carrara Formation (California, United States), tentatively assigned to the genus Discophyllum (an animal of uncertain phylogenetic placement, might be a chondrophore or an eldoniid) is published by Lieberman et al. (2017).^[237]
Specimens of Cloudina associated with microbial mat textures are reported from the Ediacaran Tamengo Formation (Brazil) by Becker-Kerber et al. (2017).^[238]
An assemblage of trace fossils from Ediacaran–Cambrian siltstones in Brazil, probably produced by a nematoid-like organism, is described by Parry et al. (2017).^[239]
A diverse fauna dominated by sponges living immediately after the Hirnantian extinction is described from China by Botting et al. (2017).^[240]
A diverse Early Triassic (Olenekian) marine assemblage (Paris biota), including leptomitid protomonaxonid sponges (a group otherwise known only from Cambrian and Ordovician), new forms of the crinoid order Holocrinida displaying advanced characters, a probable basal ophiodermatid and gladius-bearing coleoids (previously unknown in Early Triassic strata) is reported from Paris (Idaho, United States) by Brayard et al. (2017).^[241]
A study on the muscle anatomy of Pambdelurion whittingtoni is published by Young & Vinther (2017).^[242]
Cambrian species Zhenghecaris shankouensis, originally classified as a bivalved arthropod, is reinterpreted as a member of Radiodonta by Zeng et al. (2017).^[243]
The holotype specimen of a putative lobopodian species Aysheaia prolata is reinterpreted as an isolated frontal appendage of a radiodontan belonging to the genus Stanleycaris by Pates, Daley & Ortega-Hernández (2017).^[244]
A revision of the radiodontan genus Caryosyntrips is published by Pates & Daley (2017), who interpret the holotype specimen of a putative lobopodian species Mureropodia apae as a partial isolated appendage of a member of the genus Caryosyntrips.^[245]
Description of the morphology of Amplectobelua symbrachiata, with a focus on its head region, is published by Cong et al. (2017).^[246]
A study on the anatomy of the Cambrian hyolith Haplophrentis, as well as on the phylogenetic relationships of the hyoliths, is published by Moysiuk, Smith & Caron (2017).^[247]
A study on the phylogenetic relationships of Tullimonstrum gregarium, challenging its interpretation as a vertebrate, is published by Sallan et al. (2017).^[248]
New exceptionally preserved fossils of Vetulicola longbaoshanensis are described from the Lower Cambrian Wulongqing Formation (China) by Li, Liu & Ou (2017).^[249]
Putative trematode metacercariae preserved at the base of the femora of an agamid lizard are described from the Cretaceous Burmese amber (Myanmar) by Poinar et al. (2017).^[250]

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Acoelia discontinua^[251]	Sp. nov	Valid	Wu	Permian (Changhsingian)		China	A calcareous sponge belonging to the order Inozoa and the family Acoeliidae.
Aeroretiolites^[252]	Gen. et sp. nov	Valid	Melchin, Lenz & Kozłowska	Silurian		Canada	A graptolite. Genus includes new species A. cancellatus.
Aladraco^[253]	Nom. et sp. nov	Valid	Geyer	Cambrian	Jbel Wawrmast Formation Tannenknock Formation	Germany Morocco	A member of Hyolitha; a replacement name for Oxyprymna Kiderlen (1933). Genus includes A. schloppensis (Wurm, 1925) and a new species A. ougnatensis.
Allonnia erjiensis^[254]	Sp. nov	Valid	Yun, Zhang & Li	Cambrian	Chengjiang Lagerstätte	China	A chancelloriid.
Andiprion^[255]	Gen. et sp. nov	Valid	Hints et al.	Ordovician (Dapingian)		Argentina	A polychaete described on the basis of scolecodonts. Genus includes new species A. paxtonae.
Angulosuspongia^[256]^[257]	Gen. et sp. nov	Valid	Yang et al.	Cambrian Stage 5	Kaili Formation	China	A sponge belonging to the order Verongida and the family Vauxiidae. Genus includes new species A. sinensis.
Ankalodous^[258]	Gen. et sp. nov	Valid	Shu et al.	Cambrian Series 3	Qiongzhusi (Chiungchussu) Formation	China	An arrow worm. The type species is A. sericus.
Archaeochionelasmus^[259]	Gen. et sp. nov	Valid	Kočí et al.	Late Cretaceous (Cenomanian)	Bohemian Cretaceous Basin	Czech Republic	An animal of uncertain phylogenetic placement. Originally interpreted as a barnacle belonging to the group Balanomorpha and the superfamily Chionelasmatoidea; Gale & Skelton (2018) considered it to be a rudist bivalve instead.^[260] Genus includes new species A. nekvasilovae.
Biskolites^[261]	Gen. et sp. nov	Valid	Valent, Fatka & Marek	Cambrian (Drumian)	Buchava Formation	Czech Republic	A member of Hyolitha. Genus includes new species B. iactans.
Capinatator^[262]	Gen. et sp. nov	Valid	Briggs & Caron	Cambrian	Burgess Shale	Canada ( British Columbia)	An arrow worm. The type species is C. praetermissus.
Caryosyntrips camurus^[245]	Sp. nov	Valid	Pates & Daley	Cambrian	Burgess Shale Langston Formation Valdemiedes Formation?	Canada ( British Columbia) United States ( Utah) Spain?	A member of Radiodonta.
Caryosyntrips durus^[245]	Sp. nov	Valid	Pates & Daley	Cambrian	Wheeler Shale	United States ( Utah)	A member of Radiodonta.
Cloudina ningqiangensis^[263]	Sp. nov	Valid	Cai et al.	Late Ediacaran		China
Cloudina xuanjiangpingensis^[263]	Sp. nov	Valid	Cai et al.	Late Ediacaran		China
Conchicolites rossicus^[264]	Sp. nov	Valid	Vinn & Madison	Ordovician (Katian)		Russia	A member of Cornulitida belonging to the family Cornulitidae.
Conciliospongia^[265]	Gen. et sp. nov		Botting, Zhang & Muir	Late Ordovician	Wenchang Formation	China	A stem-demosponge of uncertain phylogenetic placement. The type species is C. anjiensis.
Corallistes campanensis^[266]	Sp. nov	Valid	Świerczewska-Gładysz	Late Cretaceous (early Campanian)		Poland	A lithistid demosponge belonging to the family Corallistidae.
Cretacimermis aphidophilus^[267]	Sp. nov	Valid	Poinar	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A nematode belonging to the family Mermithidae.
Eolorica^[268]	Gen. et sp. nov	Valid	Harvey & Butterfield	Cambrian (Furongian)	Deadwood Formation	Canada ( Saskatchewan)	A member of the total group of Loricifera. The type species is E. deadwoodensis.
Eorograptus spirifer^[252]	Sp. nov	Valid	Melchin, Lenz & Kozłowska	Silurian		Canada	A graptolite.
Feiyanella^[269]	Gen. et sp. nov	Valid	Han et al.	Earliest Cambrian	Kuanchuanpu Formation	China	A Cloudina-like tubular microfossil. The type species is F. manica.
Geoditesia jordaniensis^[270]	Sp. nov	Valid	Ungureanu, Ahmad & Farouk	Middle Jurassic (Callovian)		Jordan	A sponge.
Glomerula gemmellaroi^[271]	Sp. nov	Valid	Sanfilippo in Sanfilippo et al.	Permian	"Pietra di Salomone" Limestone	Italy	A polychaete belonging to the family Sabellidae, a species of Glomerula.
Guettardiscyphia zitti^[272]	Sp. nov	Valid	Vodrážka	Late Cretaceous (Turonian)	Bílá Hora Formation	Czech Republic	A hexactinellid sponge belonging to the family Cribrospongiidae.
Inquicus^[273]	Gen. et sp. nov	Valid	Cong et al.	Early Cambrian	Chengjiang Lagerstätte	China	A tiny worm infecting members of the genera Cricocosmia and Mafangscolex. Genus includes new species I. fellatus.
Keretsa^[274]	Gen. et sp. nov	Valid	Ivantsov	Late Precambrian	Zimnie Gory Formation	Russia ( Arkhangelsk Oblast)	An early eumetazoan, showing similarities to the arthropod species Naraoia longicaudata. The type species is K. brutoni.
Labechia yeongwolense^[275]	Sp. nov	Valid	Jeon et al.	Ordovician (Darriwilian)	Yeongheung Formation	South Korea	A stromatoporoid.
Lepidocoleus kuangguoduni^[276]	Sp. nov	Valid	Gügel et al.	Devonian (Eifelian)	Nandan Formation	China	A machaeridian.
'Linevitus' guizhouensis^[277]	Sp. nov	Valid	Sun et al.	Cambrian Stage 4	Balang Formation	China	A member of Hyolitha.
Microdictyon cuneum^[278]	Sp. nov	Valid	Wotte & Sundberg	Cambrian		United States ( Nevada)	A lobopodian.
Microdictyon montezumaensis^[278]	Sp. nov	Valid	Wotte & Sundberg	Cambrian		United States ( Nevada)	A lobopodian.
Mughanniyyum^[270]	Gen. et sp. nov	Valid	Ungureanu, Ahmad & Farouk	Middle Jurassic (Callovian)		Jordan	A sponge. Genus includes new species M. hanium.
Multiconotubus^[263]	Gen. et sp. nov	Valid	Cai et al.	Late Ediacaran		China	A Cloudina-like fossil. Genus includes new species M. chinensis.
Neophrissospongia kacperskii^[266]	Sp. nov	Valid	Świerczewska-Gładysz	Late Cretaceous (early Campanian)		Poland	A lithistid demosponge belonging to the family Corallistidae.
Orthrozanclus elongata^[279]	Sp. nov		Zhao & Smith in Zhao et al.	Cambrian Stage 3	Maotianshan Shales	China
Ovatiovermis^[280]	Gen. et sp. nov	Valid	Caron & Aria	Cambrian	Burgess Shale	Canada ( British Columbia)	A lobopodian belonging to the family Luolishaniidae. The type species is O. cribratus.
Pachinion canaliculatum^[266]	Sp. nov	Valid	Świerczewska-Gładysz	Late Cretaceous (early Campanian)		Poland	A lithistid demosponge belonging to the family Corallistidae.
Paratetragraptus cooperi^[281]	Sp. nov	Valid	VandenBerg	Ordovician (early Floian)		Australia	A graptolite belonging to the group Dichograptina and the family Phyllograptidae.
Paratetragraptus? henrywilliamsi^[281]	Sp. nov	Valid	VandenBerg	Ordovician (early Floian)		Australia	A graptolite belonging to the group Dichograptina and the family Phyllograptidae.
Paratetragraptus thomassmithi^[281]	Sp. nov	Valid	VandenBerg	Ordovician (early Floian)		Australia	A graptolite belonging to the group Dichograptina and the family Phyllograptidae.
Plumulites lamonti^[282]	Sp. nov	Valid	Candela & Crighton	Silurian (Telychian)	Wether Law Linn Formation	United Kingdom	A machaeridian.
Propomatoceros permianus^[271]	Sp. nov	Valid	Sanfilippo in Sanfilippo et al.	Permian	"Pietra di Salomone" Limestone	Italy	A polychaete belonging to the family Serpulidae, a species of Propomatoceros.
Pseudoretiolites hyrichus^[252]	Sp. nov	Valid	Melchin, Lenz & Kozłowska	Silurian		Canada	A graptolite.
Pyrgopolon (Septenaria) cenomanensis^[283]	Sp. nov	Valid	Kočí, Jäger & Morel	Late Cretaceous (Cenomanian)		France	A polychaete belonging to the family Serpulidae.
Pyrgopolon (Turbinia?) gaiae^[271]	Sp. nov	Valid	Sanfilippo in Sanfilippo et al.	Permian	"Pietra di Salomone" Limestone	Italy	A polychaete belonging to the family Serpulidae, a species of Pyrgopolon.
Radiofibrosclera^[251]	Gen. et sp. nov	Valid	Wu	Permian (Changhsingian)		China	A sclerosponge. The type species is R. laibinensis.
Ratcliffespongia arivechensis^[284]	Sp. nov	Valid	Beresi et al.	Cambrian Series 3		Mexico	A reticulosan sponge of uncertain phylogenetic placement.
Saccorhytus^[285]	Gen. et sp. nov	Valid	Han et al.	Earliest Cambrian		China	An animal of uncertain phylogenetic placement. Originally described as an early deuterostome related to vetulicolians and vetulocystids, but subsequently argued to be an ecdysozoan.^[286] The type species is S. coronarius.
"Serpula" distefanoi^[271]	Sp. nov	Valid	Sanfilippo in Sanfilippo et al.	Permian	"Pietra di Salomone" Limestone	Italy	A polychaete belonging to the family Serpulidae.
Serpula? pseudoserpentina^[283]	Sp. nov	Valid	Kočí, Jäger & Morel	Late Cretaceous (Cenomanian)		France	A polychaete belonging to the family Serpulidae.
Silicunculus saaqqutit^[287]	Sp. nov	Valid	Peel	Cambrian Series 3		Greenland	A sponge.
Singuuriqia^[288]	Gen. et sp. nov	Valid	Peel	Cambrian Stage 3	Sirius Passet Lagerstätte	Greenland	A member of Priapulida. Genus includes new species S. simoni.
Siphusauctum lloydguntheri^[289]	Sp. nov	Valid	Kimmig, Strotz & Lieberman	Cambrian Stage 5	Spence Shale	United States ( Utah)
Tauricornicaris^[243]	Gen. et 2 sp. nov	Valid^[290]	Zeng et al.	Early Cambrian	Chengjiang Lagerstätte	China	Originally considered as member of Radiodonta, possibly a member of Hurdiidae, but denied in 2018.^[291]^[292] Genus includes new species T. latizonae and T. oxygonae.
Thoracospongia lacrimiformis^[287]	Sp. nov	Valid	Peel	Cambrian Series 3		Greenland	A sponge.
Tianzhushanella tolli^[293]	Sp. nov	Valid	Kouchinsky et al.	Cambrian	Medvezhya Formation	Russia	A member of Tianzhushanellidae (a group of animals of uncertain phylogenetic placement, possibly stem-brachiopods).
Tshallograptus^[281]	Gen. et comb. et 3 sp. nov	Valid	VandenBerg	Ordovician (early Floian)		Australia Canada	A graptolite belonging to the group Dichograptina and the family Phyllograptidae. The type species is "Graptolithus" fruticosus Hall (1858); genus also includes new species T. tridens, T. cymulus and T. furcillatus.
Valospongia sonorensis^[284]	Sp. nov	Valid	Beresi et al.	Cambrian Series 3		Mexico	A reticulosan sponge of uncertain phylogenetic placement.
Vittatusivermis^[294]	Gen. et sp. nov		Zhang et al.	Cambrian (Fortunian)	Yuhucun Formation	China	A worm-like organism, possibly a member of Bilateria of uncertain phylogenetic placement. The type species is V. annularius.
Websteroprion^[295]	Gen. et sp. nov	Valid	Eriksson, Parry & Rudkin	Devonian (late Emsian-early Eifelian)	Kwataboahegan Formation	Canada ( Ontario)	A eunicidan polychaete of uncertain phylogenetic placement. The type species is W. armstrongi.

Other organisms

Research

Eoarchean (over 3,700 million years old) organic residues are reported from Isua, West Greenland by Hassenkam et al. (2017).^[296]
Putative fossilized microorganisms that are at least 3,770 million and possibly 4,280 million years old are described from the Nuvvuagittuq belt (Quebec, Canada) by Dodd et al. (2017).^[297]
Organic carbon contents are reported from the oldest metasedimentary rocks from northern Labrador (Canada) by Tashiro et al. (2017), who interpret the finding as the oldest evidence of organisms greater than 3.95 Ga;^[298] the study is subsequently criticized by Whitehouse et al. (2019).^[299]
Potential biosignatures, including stromatolites, are reported from the newly discovered rocks recovered from ca. 3.48 billion years old Dresser Formation (Pilbara Craton, Australia) by Djokic et al. (2017).^[300]
Lenticular structures known from the ~3.4 Ga Kromberg Formation (Kaapvaal Craton, South Africa) are interpreted as organic Archean microfossils by Oehler et al. (2017).^[301]
Fossils of early eukaryotes Tappania plana, Dictyosphaera macroreticulata and Valeria lophostriata are described from the early Mesoproterozoic Greyson Formation (Belt Supergroup, Montana, United States) by Adam et al. (2017).^[302]
2.4-billion-year-old filamentous fossils forming mycelium-like structures, considered to be either the oldest known fungi or members of an unknown branch of fungus-like mycelial organisms, are described from the Ongeluk Formation (South Africa) by Bengtson et al. (2017).^[303]
A study on the anatomy of the fossils of Chuaria circularis recovered from the Tonian Liulaobei Formation (China) is published by Tang et al. (2017), who interpret Chuaria as most likely a simple multicellular organism (a colonial organism without cell differentiation).^[304]
A study on the apatitic scale microfossils from the Fifteenmile Group (Yukon, Canada), indicating that the fossils document the existence of eukaryotic biomineralizing organisms approximately 810 million years ago, is published by Cohen et al. (2017).^[305]
A study on the structure, morphology, and development of the large intracellular structures preserved in embryo-like microfossils from the Ediacaran Weng'an Biota (China) is published by Yin et al. (2017), who interpret these structures as likely cell nuclei.^[306]
A study testing the suggested link between the appearance of large body size in rangeomorphs (organisms of uncertain phylogenetic placement, likely animals) in the Ediacaran and postulated regional increases in environmental nutrient levels is published by Hoyal Cuthill & Conway Morris (2017).^[307]
A study on the internal morphology of Rangea from the Nama Group (Namibia), based on data obtained using X-ray micro-computed tomography, is published by Sharp et al. (2017).^[308]
Smith et al. (2017) report the discovery of fossils of Gaojiashania from the Ediacaran strata of the Nama Group (Namibia) and a new fossil assemblage from the Ediacaran strata of the Wood Canyon Formation (Nevada, United States), including erniettomorphs and a variety of tubular body fossils.^[309]
A study on the well-preserved Devonian calcareous nanicellid foraminiferans from the Świętokrzyskie Mountains (Poland) and their implications for the biomineralization style and affinities of Paleozoic fusulinid foraminiferans is published by Dubicka & Gorzelak (2017).^[310]
Four forms of modern-looking gilled mushrooms, including two taxa belonging to the family Marasmiaceae, are described from the Cretaceous Burmese amber by Cai et al. (2017).^[311]

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Acadialithus^[312]	Gen. et 2 sp. nov	Valid	Howe	Late Jurassic (Tithonian)		Bulgaria Offshore eastern Newfoundland, Canada Offshore in the eastern Gulf of Mexico Offshore of the northeast coast of the United States	A nannofossil. Genus includes new species A. dennei and A. valentinei.
Adendorfia^[313]	Gen. et sp. nov	Valid	Worobiec et al.	Miocene		Germany	A fungus, probably a member of Chaetomiaceae. Genus includes new species A. miocenica.
Algites philippoviensis^[314]	Sp. nov	Valid	Naugolnykh	Permian (Kungurian)	Philippovian Formation	Russia	A brown alga.
Algites shurtanensis^[314]	Sp. nov	Valid	Naugolnykh	Permian (Kungurian)	Shurtan Formation	Russia	A brown alga.
Alpinoschwagerina nagatoensis^[315]	Sp. nov	Valid	Kobayashi	Permian (Asselian)	Akiyoshi Limestone Group	Japan	A foraminifer belonging to the group Fusulinida.
Amsassia argentina^[316]	Sp. nov	Valid	Carrera, Astini & Gomez	Early Ordovician	La Silla Formation	Argentina	A coral-like organism of uncertain phylogenetic placement.
Asterina indodeightonii^[317]	Sp. nov	Valid	Vishnu et al.	Mid-Miocene to early Pleistocene		India	A fungus, a species of Asterina.
Asterina mioconsobrina^[317]	Sp. nov	Valid	Vishnu et al.	Mid-Miocene to early Pleistocene		India	A fungus, a species of Asterina.
Asterina miosphaerelloides^[317]	Sp. nov	Valid	Vishnu et al.	Mid-Miocene to early Pleistocene		India	A fungus, a species of Asterina.
Asterina neocombreticola^[317]	Sp. nov	Valid	Vishnu et al.	Mid-Miocene to early Pleistocene		India	A fungus, a species of Asterina.
Asterina neoelaeocarpi^[317]	Sp. nov	Valid	Vishnu et al.	Mid-Miocene to early Pleistocene		India	A fungus, a species of Asterina.
Asterina presaracae^[317]	Sp. nov	Valid	Vishnu et al.	Mid-Miocene to early Pleistocene		India	A fungus, a species of Asterina.
Baculogypsinella^[318]	Gen. et sp. nov	Valid	Matsumaru	Eocene		Philippines	A foraminifer. Genus includes new species B. eocenica.
Blastanosphaira^[319]	Gen. et sp. nov	Valid	Javaux & Knoll	Mesoproterozoic	Mainoru Formation	Australia	A possible eukaryotic microorganism of uncertain phylogenetic placement. The type species is B. kokkoda.
Bonniea makrokurtos^[320]	Sp. nov	Valid	Cohen, Irvine & Strauss	Tonian	Callison Lake Formation	Canada ( Yukon)	A vase-shaped microfossil.
Braarudosphaera pseudobatilliformis^[321]	Sp. nov	Valid	Alves, Lima & Shimabukuro	Early Cretaceous (Aptian)		Brazil	A haptophyte belonging to the family Braarudosphaeraceae.
Carbonoschwagerina nipponica^[315]	Sp. nov	Valid	Kobayashi	Carboniferous (Kasimovian and Gzhelian)	Akiyoshi Limestone Group	Japan	A foraminifer belonging to the group Fusulinida.
Cephalothecoidomyces^[313]	Gen. et sp. nov	Valid	Worobiec et al.	Neogene		Germany Poland	A fungus, probably a member of Cephalothecaceae. Genus includes new species C. neogenicus.
Chiphragmalithus muzylevii^[322]	Sp. nov	Valid	Musatov	Eocene (Ypresian)		Russia	A haptophyte.
Cobios^[323]	Gen. et sp. nov	Valid	Du et al.	Ediacaran	Doushantuo Formation	China	A red alga. The type species is Cobios rubo.
Curviacus^[324]	Gen. et sp. nov	Valid	Shen et al.	Ediacaran	Dengying Formation	China	A benthic modular organism consisting of serially arranged and crescent-shaped chambers. Genus includes new species C. ediacaranus.
Cyanonema grandis^[325]	Sp. nov	Valid	Shi & Feng in Shi et al.	Early Mesoproterozoic	Gaoyuzhuang Formation	China	A member of Cyanobacteria belonging to the group Nostocales.
Cycliocyrillium rootsi^[320]	Sp. nov	Valid	Cohen, Irvine & Strauss	Tonian	Callison Lake Formation Chuar Group (Kwagunt Formation)^[326]	Canada ( Yukon) United States^[326]	A vase-shaped microfossil. Originally described as a species of Cycliocyrillium, but subsequently transferred to the genus Obelix.^[326] Morais et al. (2019) corrected the suffix for the specific epithet to rootsii.^[326]
Dalongicaepa^[327]	Gen. et sp. et comb. nov	Valid	Xiao & Suzuki in Xiao, Suzuki & He	Late Permian	Upper Dalong Formation	China Thailand	A radiolarian belonging to the group Spumellaria and the family Spongotortilispinidae. The type species is D. bipolaris; genus also includes "Pseudospongoprunum" fontainei Sashida in Sashida et al. (2000).
Denaricion^[328]	Gen. et sp. nov	Valid	Bengtson in Bengtson et al.	~1.6 billion years ago		India	An organism of uncertain phylogenetic placement, might be an alga or prokaryote. Genus includes new species D. mendax.
Devisphaera^[329]	Gen. et sp. nov	Valid	Tang et al.	Late Mesoproterozoic – early Neoproterozoic	Madhubani Group	India	An organic-walled microfossil. Genus includes new species D. corallis.
Discusphyton^[330]	Gen. et sp. nov	Valid	Wang, Wang & Du	Ediacaran	Doushantuo Formation	China	A macroalga of uncertain phylogenetic placement. Genus includes new species D. whenghuiensis.
Fissumella^[331]	Gen. et sp. nov	Valid	Cruz-Abad et al.	Early Cretaceous (Albian)		Italy	A foraminifer. Genus includes new species F. motolae.
Flabelloperforata^[332]	Gen. et sp. nov	Valid	Schlagintweit & Rashidi	Late Cretaceous (Maastrichtian)	Tarbur Formation	Iran	A foraminifer belonging to the group Loftusiida, possibly a member of the family Biokovinidae. Genus includes new species F. tarburensis.
Gigadiacrodium^[333]	Gen. et comb. et sp. nov	Valid	Szczepanik, Servais & Żylińska	Cambrian (Furongian)	Alum Shale Formation Elliott's Cove Formation	Canada Iran Italy Poland Sweden	An acritarch. The type species is "Veryhachium" martinum Pittau (1985); genus also includes new species G. vidalii.
Gigantosphaeridium floccosum^[334]	Sp. nov	Valid	Agić, Moczydłowska & Yin	Early Mesoproterozoic	Ruyang Group	China	A microfossil.
Gondwanagaricites^[335]^[336]	Gen. et sp. nov	Valid	Heads, Miller & Crane	Early Cretaceous (Aptian)	Crato Formation	Brazil	A gilled mushroom. Genus includes new species G. magnificus.
Hagenococcus^[337]	Gen. et sp. nov	Valid	Krings et al.	Early Devonian	Rhynie chert	United Kingdom	A microorganism of uncertain phylogenetic placement, most likely an alga with affinities to the Chlorophyta or Streptophyta. Genus includes new species H. aggregatus.
Haplophragmoides arcticus^[338]	Sp. nov	Valid	Kaminski, Waskowska & Chan	Middle Pleistocene		Arctic Ocean (Lomonosov Ridge)	A foraminifer.
Jigulites titanicus^[315]	Sp. nov	Valid	Kobayashi	Carboniferous (Gzhelian) and Permian (Asselian)	Akiyoshi Limestone Group	Japan	A foraminifer belonging to the group Fusulinida.
Limeta^[339]	Gen. et sp. nov	Valid	Morais, Fairchild & Lahr in Morais et al.	Neoproterozoic	Urucum Formation	Brazil	A vase-shaped microfossil. Genus includes new species L. lageniformis.
Montiparus minensis^[315]	Sp. nov	Valid	Kobayashi	Carboniferous (Kasimovian)	Akiyoshi Limestone Group	Japan	A foraminifer belonging to the group Fusulinida.
Nannoconus troelsenii^[321]	Sp. nov	Valid	Alves, Lima & Shimabukuro	Early Cretaceous (Aptian)		Brazil	A haptophyte belonging to the family Nannoconaceae.
Oscillatoriopsis gigas^[325]	Sp. nov	Valid	Shi & Feng in Shi et al.	Early Mesoproterozoic	Gaoyuzhuang Formation	China	A member of Cyanobacteria belonging to the group Oscillatoriales.
Palaeoamphora^[339]	Gen. et sp. nov	Valid	Morais, Fairchild & Lahr in Morais et al.	Neoproterozoic	Urucum Formation	Brazil	A vase-shaped microfossil. Genus includes new species P. urucumense.
Palaeostromatus^[340]	Gen. et sp. nov	Valid	Dentzien-Dias, Poinar & Francischini	Permian (Guadalupian)	Rio do Rasto Formation	Brazil	An actinomycete. Genus includes new species P. diairetus.
Paleohaimatus^[341]	Gen. et sp. nov	Valid	Poinar	Eocene-Miocene	El Mamey Formation (Dominican amber)	Dominican Republic	A member of Apicomplexa belonging to the group Piroplasmida. Genus includes new species P. calabresi.
Parastaffelloides kanmerai^[315]	Sp. nov	Valid	Kobayashi	Carboniferous (Moscovian)	Akiyoshi Limestone Group	Japan	A foraminifer belonging to the group Fusulinida.
Pentadinium darmirae^[342]	Sp. nov	Valid	Slimani & Ţabără in Ţabără et al.	Paleocene (Danian)	Izvor Formation Runcu Formation	Romania	A dinoflagellate belonging to the group Gonyaulacales and the family Gonyaulacaceae.
Persiella^[332]	Gen. et sp. nov	Valid	Schlagintweit & Rashidi	Late Cretaceous (Maastrichtian)	Tarbur Formation	Iran	A foraminifer belonging to the group Loftusiida, possibly a member of the family Spirocyclinidae. Genus includes new species P. pseudolituus.
Pocillithus crucifer^[343]	Sp. nov	Valid	Lees, Bown & Young	Late Cretaceous (Turonian)		Tanzania	A haptophyte belonging to the family Papposphaeraceae.
Pocillithus macleodii^[343]	Sp. nov	Valid	Lees, Bown & Young	Late Cretaceous (Turonian)		Tanzania	A haptophyte belonging to the family Papposphaeraceae.
Quasifusulinoides grandis^[315]	Sp. nov	Valid	Kobayashi	Carboniferous (Kasimovian)	Akiyoshi Limestone Group	Japan	A foraminifer belonging to the group Fusulinida.
Rafatazmia^[328]	Gen. et sp. nov	Valid	Bengtson in Bengtson et al.	~1.6 billion years ago		India	An alga of uncertain phylogenetic placement. Genus includes new species R. chitrakootensis.
Ramathallus^[328]	Gen. et sp. nov	Valid	Sallstedt in Bengtson et al.	~1.6 billion years ago		India	A possible stem-florideophycean red algae. Genus includes new species R. lobatus.
Schwagerina wakatakeyamensis^[315]	Sp. nov	Valid	Kobayashi	Permian (Asselian)	Akiyoshi Limestone Group	Japan	A foraminifer belonging to the group Fusulinida.
Schwagerina watanabei^[315]	Sp. nov	Valid	Kobayashi	Permian (Asselian)	Akiyoshi Limestone Group	Japan	A foraminifer belonging to the group Fusulinida.
Spearlithus^[344]	Gen. et 12 sp. nov	Valid	Da Gama	Pleistocene		Dominican Republic	A calcareous nannofossil of uncertain phylogenetic placement.
Staffella subsphaerica^[315]	Sp. nov	Valid	Kobayashi	Carboniferous (Kasimovian and Gzhelian)	Akiyoshi Limestone Group	Japan	A foraminifer belonging to the group Fusulinida.
Stradnerlithus? haynesiae^[343]	Sp. nov	Valid	Lees, Bown & Young	Late Cretaceous (Turonian)		Tanzania	A haptophyte belonging to the order Stephanolithiales and the family Stephanolithiaceae.
Stradnerlithus wendleri^[343]	Sp. nov	Valid	Lees, Bown & Young	Late Cretaceous (Turonian)		Tanzania	A haptophyte belonging to the order Stephanolithiales and the family Stephanolithiaceae.
Suraqalatia^[345]	Gen. et sp. nov	Valid	Görmüş, Ameen Lawa & Al Nuaimy	Late Cretaceous (Maastrichtian)		Iraq	A foraminifer belonging to the family Dicyclinidae. Genus includes new species S. brasieri.
Synaptomitus^[346]	Gen. et sp. nov	Valid	Poinar	Eocene to Miocene	Dominican amber	Dominican Republic	Originally described as a fungus belonging to the group Basidiomycota,^[346] but this interpretation was challenged by Selosse et al. (2017).^[347] Genus includes new species S. orchiphilus.
Syracosphaera antiqua^[343]	Sp. nov	Valid	Bown, Lees & Young	Late Cretaceous (Turonian)		Tanzania	A haptophyte belonging to the order Syracosphaerales and the family Syracosphaeraceae.
Syracosphaera repagula^[343]	Sp. nov	Valid	Bown, Lees & Young	Late Cretaceous (Turonian)		Tanzania	A haptophyte belonging to the order Syracosphaerales and the family Syracosphaeraceae.
Tarburina^[348]	Gen. et sp. nov	Valid	Schlagintweit, Rashidi & Barani	Late Cretaceous (late Maastrichtian)	Tarbur Formation	Iran	A foraminifer. Genus includes new species T. zagrosiana.
Taruma^[339]	Gen. et sp. nov	Valid	Morais, Fairchild & Lahr in Morais et al.	Neoproterozoic	Urucum Formation	Brazil	A vase-shaped microfossil. Genus includes new species T. rata.
Tortolithus foramen^[343]	Sp. nov	Valid	Lees, Bown & Young	Late Cretaceous (Turonian)		Tanzania	A haptophyte of uncertain phylogenetic placement.
Veteronostocale grandis^[325]	Sp. nov	Valid	Shi & Feng in Shi et al.	Early Mesoproterozoic	Gaoyuzhuang Formation	China	A member of Cyanobacteria belonging to the group Nostocales.
Windipila^[349]	Gen. et sp. nov	Valid	Krings & Harper	Early Devonian	Windyfield chert	United Kingdom	A fungus described on the basis of a reproductive unit. Genus includes new species W. spinifera.
Xiaohongyuia^[350]	Gen. et sp. nov	Valid	Shi & Feng in Shi et al.	Late Paleoproterozoic	Dahongyu Formation	China	A probable eukaryotic microfossil. Genus includes new species X. sinica.

General paleontology

Research related to paleontology that either does not concern any of the groups of the organisms listed above, or concerns multiple groups.

A study on the links between changes in the composition of exposed continental crust and oxygenation of the atmosphere in the Precambrian is published by Smit & Mezger (2017).^[351]
A review of the progress in modeling the Snowball Earth atmosphere, cryosphere, hydrosphere and lithosphere, specifically as it pertains to Cryogenian geology and geobiology, is published by Hoffman et al. (2017).^[352]
A revised record of fossil eukaryotic steroids during the Neoproterozoic is presented by Brocks et al. (2017), who argue that bacteria were the only notable primary producers in the oceans before the Cryogenian, and that rapid rise of marine planktonic algae to domination occurred in the narrow time interval between the Sturtian and Marinoan glaciations, 659–645 million years ago, likely driving the subsequent radiation of animals in the Ediacaran period.^[353]
A study evaluating whether mass extinction events over the last 500 million year were caused by astronomical phenomena is published by Erlykin et al. (2017).^[354]
A study on the water column geochemistry of the Yangtze Sea during the Ediacaran-Cambrian transition and its implications for the relationship between ocean oxygenation and Early Cambrian animal diversification is published by Zhang et al. (2017).^[355]
A study on the links between the expansion of siliceous sponges and seawater oxygenation during the Ediacaran–Cambrian transition is published by Tatzel et al. (2017).^[356]
A study on the factors influencing marine invertebrate diversity dynamics through the Phanerozoic is published by Cermeño et al. (2017).^[357]
Edwards et al. (2017) identify a strong temporal link between the rising atmospheric oxygen levels and the Great Ordovician Biodiversification Event.^[358]
A study on the impact of the drawdown of atmospheric carbon dioxide (caused by burial of organic carbon leading to the formation of coal) on the climate around the Carboniferous/Permian boundary is published by Feulner (2017).^[359]
A comprehensive reconstruction of the Permian (Lopingian) Bletterbach Biota (Italy) and a review of other best-known Lopingian terrestrial associations containing both vertebrate and plant remains is published by Bernardi et al. (2017).^[360]
A study on the causal connection between the Siberian Traps large igneous province magmatism and Permian–Triassic extinction event, identifying the initial emplacement pulse as likely to have triggered mass extinction, is published by Burgess, Muirhead & Bowring (2017).^[361]
Viglietti, Rubidge & Smith (2017) review the tectonic setting of the Late Permian Karoo Basin (South Africa), provide an updated basin development model, and interpret their findings as indicating that the climatic changes associated with the Permian–Triassic extinction event were occurring much lower in the stratigraphy (and thus earlier) than previously documented.^[362]
A summary of knowledge of the impact of Permian-Triassic mass extinction on reef ecosystems, and on their recovery after this extinction, is presented by Martindale, Foster & Velledits (2017).^[363]
A study on benthic invertebrate communities from the Lower Triassic Werfen Formation (Italy), aiming to test whether carbon isotope perturbations during the Early Triassic were associated with biotic crises that impeded benthic recovery after the Permian–Triassic extinction event, is published by Foster et al. (2017).^[364]
A study on the impact of the magmatic activity associated with the Central Atlantic magmatic province on the Triassic–Jurassic extinction event is published by Davies et al. (2017).^[365]
A study on the volcanic activity at the end of the Triassic as indicated by mercury concentrations in sediments from around the world is published by Percival et al. (2017).^[366]
A study on the oxygen levels in Earth's oceans during and after the Triassic–Jurassic extinction event as indicated by uranium isotopes in shallow-marine limestones in the Lombardy Basin (northern Italy) is published by Jost et al. (2017).^[367]
A high-resolution stratigraphic chart for terrestrial Late Cretaceous units of North America and a study on the stratigraphic ranges of North American dinosaurs is published by Fowler (2017).^[368]
A study on the impact that large amounts of soot injected into the atmosphere during the Cretaceous–Paleogene extinction event (probably caused by global wildfires) had on the climate is published by Bardeen et al. (2017).^[369]
A study estimating the decrease of the air temperature and the duration of the climate cooling caused by Chicxulub impact at the end of the Cretaceous is published by Brugger, Feulner & Petri (2017).^[370]
A study on the volume of the climate-active gases released from sedimentary rocks as a result of the Chicxulub impact, as well as on their effect on the global climate, is published by Artemieva, Morgan & Expedition 364 Science Party (2017).^[371]
Kaiho & Oshima (2017) calculate the amounts of stratospheric soot and sulfate formed by a virtual asteroid impact at various global locations, and conclude that the Cretaceous–Paleogene extinction event was caused by the Chicxulub impact happening at the hydrocarbon-rich, sulfate-dominated area on the Earth's surface, and that an impact at a low–medium hydrocarbon area on Earth would be unlikely to cause mass extinction.^[372]
A study on the data sets of molluscan fossils from the Cretaceous–Paleogene of the Seymour Island (Antarctica) is published by Tobin (2017), who identifies possible evidence of two separate extinction events, one prior to the Cretaceous–Paleogene boundary, and one simultaneous with the bolide impact at the Cretaceous–Paleogene boundary.^[373]
A study on the behavioral and ecological diversification of animals that colonized land as indicated by trace fossils is published by Minter et al. (2017).^[374]
A study on the age of the Cowie Harbour Fish Bed (Scotland, United Kingdom), containing fish and arthropod fossils (including the millipede Pneumodesmus newmani), is published by Suarez et al. (2017).^[375]
A study on the preservation of skin and keratinous integumentary structures in tetrapod fossils through time is published by Eliason et al. (2017).^[376]
A study on the differences between the tetrapod faunas at different latitudes during the early and middle Permian, as well as their implications for establishing whether the Olson's Extinction was a genuine event, is published by Brocklehurst et al. (2017).^[377]
A study on the non-flying terrestrial tetrapod species richness through the Mesozoic and early Palaeogene is published by Close et al. (2017).^[378]
A study on the evolution of the shape of brain and skull roof during the transition from early reptiles through archosauromorphs, including nonavian dinosaurs, to birds is published by Fabbri et al. (2017).^[379]
A study on the structure and vulnerability of the food web in marine vertebrate assemblages prior to the Cretaceous–Paleogene extinction event as indicated by calcium isotope data from plesiosaurs and mosasaurs is published by Martin et al. (2017).^[380]
Qvarnström et al. (2017) reconstruct fossil inclusions in two coprolites (produced by an insectivorous animal and a large aquatic predator) from the Late Triassic locality of Krasiejów (Poland) using propagation phase-contrast synchrotron microtomography.^[381]
A study on the fossil inclusions in coprolite fragments (produced by medium to large-sized carnivores, possibly therocephalian therapsids or early archosauriforms) recovered from the Late Permian locality of Vyazniki (Russia) is published by Bajdek et al. (2017).^[382]
A new tetrapod assemblage from the lowermost levels of the Triassic Chañares Formation (Argentina), dominated by fossils of Tarjadia ruthae, dicynodonts and cynodonts, and also including fossils of other pseudosuchians and rhynchosaurs, is described by Ezcurra et al. (2017), who also reinterpret Tarjadia ruthae and Archeopelta arborensis as erpetosuchid archosaurs.^[383]
A study on the cosmopolitanism of terrestrial amniote faunas in the aftermath of the Permian–Triassic extinction event and Triassic–Jurassic extinction event is published by Button et al. (2017).^[384]
Frese et al. (2017) determine the mineral and elemental composition of a range of fossils from the Talbragar fossil site (Australia) and their rock matrices using ultraviolet light-induced fluorescence/photoluminescence, X-ray fluorescence and X-ray diffractometry, and use those techniques to reveal anatomical details of animals and plants fossils that weren't discernible otherwise.^[385]
A study on changes of the size of fossil marine shells and predatory drill holes in those shells during the Phanerozoic, as well as their implications for changes of predator-prey size ratio throughout the Phanerozoic, is published by Klompmaker et al. (2017).^[386]
A study evaluating the utility of oxygen-isotope compositions of fossilised foraminifera tests as proxies for surface- and deep-ocean paleotemperatures, and its implications for inferring Late Cretaceous and Paleogene deep-ocean and high-latitude surface-ocean temperatures, is published by Bernard et al. (2017).^[387]^[388]^[389]
A study on the glacial development and environmental changes in the Aurora Subglacial Basin (Antarctica) throughout the Cenozoic based on geophysical and geological evidence is published by Gulick et al. (2017).^[390]
A study on the onset duration of the Paleocene–Eocene Thermal Maximum is published by Kirtland Turner et al. (2017).^[391]
A study on the relationship between volcanic activity in the North Atlantic Igneous Province and the Paleocene–Eocene Thermal Maximum is published by Gutjahr et al. (2017).^[392]
A study on the environment in the area corresponding to the present-day Amazon basin in the Miocene as indicated by data from the shark and ray fossils from the Pirabas Formation (Brazil) is published by Aguilera et al. (2017).^[393]
A study on the impact of the Messinian salinity crisis on Mediterranean magmatism is published by Sternai et al. (2017).^[394]
A study on the changes of ice sheets volume and sea level during the late Pliocene is published by de Boer et al. (2017).^[395]
Pimiento et al. (2017) identify a previously unrecognized extinction event among marine megafauna at the end of the Pliocene.^[396]
A study on the aridity in eastern Africa over the past 4.4 million years as indicated by oxygen isotope ratios in fossil herbivore tooth enamel, and on its implications for inferring the role of climate in shaping early hominin environments, is published by Blumenthal et al. (2017).^[397]
Tierney, deMenocal & Zander (2017) reconstruct temperature and aridity in the Horn of Africa region spanning the past 200,000 years.^[398]
A vertebrate fauna from the Pleistocene and Holocene of Sumba (Indonesia) is described by Turvey et al. (2017).^[399]
A study on the modified mammalian bones from the Plio–Pleistocene of Ethiopia is published by Sahle, El Zaatari & White (2017), who interpret the marks on some of these bones as more likely to be produced by crocodiles than by hominids using stone tools.^[400]
Hagstrum et al. (2017) report impact-related microspherules and elevated platinum concentrations found in fine-grained sediments retained within Late Pleistocene bison and mammoth skull fragments from Alaska and Yukon, and interpret the findings as evidence of repeated airbursts and ground/ice impacts associated with multiple episodes of cosmic impact.^[401]
A study on changes in landscape moisture in the rangelands in Europe, Siberia and the Americas during the late Pleistocene as indicated by data from the bones of megaherbivores is published by Rabanus-Wallace et al. (2017).^[402]

Flora

Cnidarians

Research

New taxa

Arthropods

Bryozoans

Research

New taxa

Brachiopods

Research

New taxa

Molluscs

Echinoderms

Research

New taxa

Conodonts

Research

New taxa

Fishes

Amphibians

Research

New taxa

Temnospondyls

Lissamphibians

Other amphibians

Reptiles

Synapsids

Non-mammalian synapsids

Research

New taxa

Mammals

Other animals

Research

New taxa

Other organisms

Research

New taxa

General paleontology

References