Shepherd tree
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Eudicots
Clade:	Rosids
Order:	Brassicales
Family:	Capparaceae
Genus:	Boscia
Species:	B. albitrunca
Binomial name
Boscia albitrunca (Burch.) Gilg & Ben.

Boscia albitrunca, commonly known as the shepherd tree or shepherd's tree (Afrikaans: Witgat, Sotho: Mohlôpi, Tswana: Motlôpi, Venda: Muvhombwe, Xhosa: Umgqomogqomo, Zulu: Umvithi), is a protected species of South African tree in the caper family.^[1] It is known for having the deepest known root structure of any plant at 68 metres (223 ft).^[2]

The species epithet "albitrunca" refers to the white trunk it oftentimes develops. Traditionally, the shepherd tree was used by Dutch settlers, boers, to create a variant of coffee from the roots of the tree.^[3] It is an evergreen tree native to southern and tropical Africa, living in the hot, dry, and often seasonally brackish low-lying areas, sometimes on abundant lime or occasionally on rocky terrain. It is a common tree of the Kalahari, bushveld and lowveld. It is one of the most important animal forage trees in the Kalahari.^[4]

Description

This tree grows up to 10 m (33 ft) tall but is usually much smaller. It has a prominent, sturdy white trunk frequently with strips of rough, dark-coloured bark. The crown is often browsed by antelope and any grazers capable of reaching the foliage, resulting in a conspicuous flattened underside, or browse-line. The leaves are narrow, oblanceolate, and stiff, with veins obscure except for the distinct midrib. The flowers are small, greenish-yellow, lacking petals, starburst-shaped, and clustered. The fruits, on a jointed stalk, are about 10 mm (0.4 in) in diameter and are brittle-skinned with a whitish flesh and large endocarp. A specimen found in the central Kalahari in 1974 had roots extending to 68 m (223 ft) deep, making it the plant with the deepest known roots thus far found.^[2]

Relationships

Boscia belongs to the caper family, Capparaceae. Boscia albitrunca is closely related to Boscia foetida subsp. rehmanniana, the bushveld shepherd's tree, which has much smaller leaves and velvet-textured skin on its fruits. The genus was named for Louis Bosc (1759–1828), a French professor of agriculture who lived through the French Revolution.^{[citation needed]}

Gallery

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  Specimen with a clear browse line, Limpopo

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  Specimen with its canopy infested with red-berry mistletoe, Limpopo

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  Flowering during late winter, Limpopo

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  Green fruit during spring, Kalahari

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  Ripe fruit consumed by a cape starling, early summer, Kgalagadi