Staphylea colchica
Scientific classification Edit this classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Rosids
Clade: Malvids
Order: Crossosomatales
Takht. ex Reveal[1]

The Crossosomatales are an order, first recognized as such by APG II. They are flowering plants included within the Rosid eudicots.


Species assigned to the Crossosomatales have in common flowers that are positioned solitarily, with the base of the calyx, corolla, and stamens fused into a tube-shaped floral cup, sepals overlapping, the outermost smaller than the inner. Insides of the casings of pollen grains have horizontally extended thin regions (or endo-apertures). The gynoecium is placed on a short stalk, papillae on the stigma consist of two or more cells, ovary locules taper upwards, and the protective cell layer (or integument) surrounding the ovule leaves a zigzag opening (or micropyle). Some cell clusters have bundles of long yellow crystals, mucilage cells are present, and seeds have a smooth, woody coating.[2]


The relationships between orders within the Malvid clade, according to the APG system, is represented by the following tree.[3]


Within the Crossosomatales, the APG III system of 2009 recognises families represented in the following tree.[4]



  1. ^ Angiosperm Phylogeny Group (2009). "An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG III". Botanical Journal of the Linnean Society. 161 (2): 105–121. doi:10.1111/j.1095-8339.2009.00996.x. hdl:10654/18083.
  2. ^ Matthews M.R, Endress P.K. (2005). "Comparative floral structure and systematics in Crossosomatales (Crossosomataceae, Stachyuraceae, Staphyleaceae, Aphloiaceae, Geissolomataceae, Ixerbaceae, Strasburgeriaceae)". Botanical Journal of the Linnean Society. 147 (1): 1–46. doi:10.1111/j.1095-8339.2005.00347.x.
  3. ^ Peter F. Stevens (2001). "Angiosperm Phylogeny Website".
  4. ^ Oh, S.H. (2010). "Phylogeny and systematics of Crossosomatales as inferred from chloroplast atpB, matK, and rbcL sequences". Korean Journal of Plant Taxonomy. 40 (4): 208–217. doi:10.11110/kjpt.2010.40.4.208.