Group of insects
The cohort Polyneoptera is one of the major groups of winged insects, comprising the Orthoptera (grasshoppers, crickets, etc.) and all other neopteran insects believed to be more closely related to Orthoptera than to any other insect orders. They were formerly grouped together with the Palaeoptera and Paraneoptera as the Hemimetabola or Exopterygota on the grounds that they have no metamorphosis, the wings gradually developing externally throughout the nymphal stages.[2] Many members of the group have leathery forewings (tegmina) and hindwings with an enlarged anal field (vannus).
Taxonomy
Extant
The following extant orders are included in Polyneoptera:[3]
Fossil
The following fossil groups are included in Polyneoptera:[3]
- Archaeorthoptera (Orthoptera and stem-group relatives)
- †Caloneurodea
- †Cnemidolestodea (= Cnemidolestida)[4][5]
- †Geraroptera
- †Titanoptera – Carboniferous to Triassic
- order Incertae sedis
- family †Cacurgidae Handlirsch, 1911
- family †Chresmodidae Haase, 1890[6]
- family †Permostridulidae Béthoux, Nel, Lapeyrie & Gand, 2003
- family †Protophasmatidae Brongniart, 1885
- genus †Chenxiella Liu, Ren & Prokop, 2009
- genus †Lobeatta Béthoux, 2005
- genus †Longzhua Gu, Béthoux & Ren, 2011
- genus †Nectoptilus Béthoux, 2005
- genus †Sinopteron Prokop & Ren, 2007
- Stem-group Phasmatodea
- †family Xiphopteridae Sharov 1968
- †family Prochresmodidae Vishnyakova 1980
- †family Aeroplanidae Tillyard 1918
- †family Cretophasmatidae Sharov 1968
- †family Aerophasmatidae Martynov, 1928
- Stem-group Dermaptera
- †"Grylloblattida" (Stem-group Grylloblattodea?)
- †Eoblattida
- †Paoliida[7]
- †Protorthoptera
- † family Magicivenidae[8]
Phylogeny
The following cladogram is based on the molecular phylogeny of Wipfler et al. 2019:[9]
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