Proteles Temporal range: Late Pliocene – Recent (Possible Miocene record)

Aardwolf (Proteles cristatus)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Carnivora
Suborder:	Feliformia
Family:	Hyaenidae
Subfamily:	Protelinae Saint-Hilaire, 1851
Genus:	Proteles Saint-Hilaire, 1824
Type species
Viverra cristata Sparrman, 1783
Species
Proteles cristata †Proteles amplidentus

Proteles is a genus of distinctive hyenas which contain the aardwolf (Proteles cristatus) and its close fossil relatives.^[1] It is the only extant genus of the subfamily Protelinae.

While the oldest fossils definitely belonging to Proteles date back to the Pliocene, material from the Miocene dating to around 10 million years ago has been suggested to belong to the genus, which would significantly increase its temporal range.^[2]

It has been suggested that the Protelinae subfamily may actually be an offshoot of the "running hyenas" (such as Lycyaena) who adapted to an insectivorous diet due to increased competition from canines and felines.^[3]

References

^ Kingdon, Jonathan (2014). Mammals of Africa: Volume V: Carnivores, Pangolins, Equids and Rhinoceroses. Bloomsbury Publishing. p. 282. ISBN 9781408189948.
^ Paleontology and Geology of Laetoli: Human Evolution in Context Volume 2: Fossil Hominins and the Associated Fauna. Springer Netherlands. 2011. p. 220. ISBN 9789048199624.
^ Koepfli, Klaus-Peter (2006). "Molecular systematics of the Hyaenidae: relationships of a relictual lineage resolved by a molecular supermatrix". Molecular Phylogenetics and Evolution. 38 (3): 603–620. doi:10.1016/j.ympev.2005.10.017.

Genera of civets, mongooses, hyenas, cats, and their extinct allies

Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora

Feliformia

†Palaeogalidae

†Palaeogale?

†Nimravidae

- See Nimravidae

Aeluroidea

†Alagtsavbaatar
†Anictis
†Asiavorator
†Shandgolictis

Nandiniidae

Nandinia

Viverroidea

Viverridae

†Africanictis †Dunictis †Forsythictis †Lufengictis †Ketketictis †Mioprionodon †Moghradictis †Progenetta
Genettinae	Genetta Poiana
Hemigalinae	Chrotogale Cynogale Diplogale Hemigalus Macrogalidia
Paradoxurinae	Arctictis Arctogalidia †Kanuites †Kichechia Paguma Paradoxurus †Siamictis †Tugenictis
Viverrinae	Civettictis †Semigenetta Viverra Viverricula

Herpestoidea

Herpestidae	See below↓
Hyaenidae	See below↓

Feloidea

- See below↓

Herpestidae

Eupleridae

Euplerinae	Cryptoprocta Eupleres Fossa
Galidiinae	Galidia Galidictis Mungotictis Salanoia

Herpestidae

†Kichechia †Legetetia †Leptoplesictis †Ugandictis
Herpestinae	Atilax Bdeogale Cynictis Herpestes Ichneumia Paracynictis Urva Xenogale
Mungotinae	Crossarchus Dologale Helogale Liberiictis Mungos Suricata

Hyaenidae

†Percrocutidae?

Dinocrocuta
Percrocuta

†Lophocyonidae

Euboictis
Izmirictis
Lophocyon
Sivanasua

Hyaenidae

†Allohyaena †Belbus †Metahyaena †Pliocrocuta †Tongxinictis †Werdelinus
†Ictitheriinae	Herpestides Hyaenotherium Ictitherium Lycyaena Miohyaenotherium Plioviverrops Protictitherium Thalassictis Tungurictis
Protelinae	†Gansuyaena Proteles
Hyaeninae	†Adcrocuta †Chasmaporthetes Crocuta Hyaena †Hyaenictis †Ikelohyaena †Leecyaena †Lycyaenops †Pachycrocuta †Palinhyaena Parahyaena

Feloidea

†Pseudictis
†Stenogale
†Stenoplesictis
†Viretictis

†Barbourofelidae

(See Barbourofelidae)

Prionodontidae

†Haplogale
†Palaeoprionodon
Prionodon

Felidae

†Asilifelis †Diamantofelis †Hyperailurictis †Katifelis †Namafelis †Miopanthera †Pseudaelurus †Sivaelurus †Styriofelis
†Proailurinae	Proailurus Vinayakia
Felinae	Acinonyx Catopuma Caracal Felis Herpailurus Leopardus Leptailurus †Leptofelis Lynx †Miracinonyx Otocolobus Pardofelis †Pratifelis Prionailurus †Pristifelis Puma †Sivapanthera †Sivapardus †Vishnufelis
†Machairodontinae	See Machairodontinae
Pantherinae	Neofelis Panthera †Leontoceryx

Taxon identifiers	Wikidata: Q18510844 Wikispecies: Proteles BOLD: 359711 CoL: 6XF9 EoL: 93385 Fossilworks: 103250 GBIF: 2433501 iNaturalist: 41891 IRMNG: 1435778 ITIS: 621800 MSW: 14000688 NCBI: 9679