Squaloziphius Temporal range: Early Miocene, 23–20 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Artiodactyla
Infraorder:	Cetacea
Family:	†Squaloziphiidae
Genus:	†Squaloziphius Muizon, 1991
Species:	†S. emlongi
Binomial name
†Squaloziphius emlongi Muizon, 1991

Squaloziphius is an extinct genus of odontocete cetacean from the Early Miocene (Aquitanian) aged marine deposits in Washington state.

Systematics

It was originally classified as the most primitive beaked whale, being placed in a separate subfamily, Squaloziphiinae,^[1] (followed by Fordyce and Muizon 2001^[2]) but later authors have placed it outside Ziphiidae as either Odontoceti incertae sedis or closely related to Ziphiidae.^[3]^[4] The description of the archaic odontocete Yaquinacetus demonstrated that Squaloziphius was by no means part of Ziphiidae and that these two taxa are more primitive than crown Odontoceti, necessitating elevation of Squaloziphiinae to full familial status, as Squaloziphiidae.^[5]

References

^ C. Muizon. 1991. A new Ziphiidae (Cetacea) from the Early Miocene of Washington State (USA) and phylogenetic analysis of the major groups of odontocetes. Bulletin du Museum National d'Histoire Naturelle. 4e Serie. Section C. Sciences de la Terre. Paleontologie, Geologie, Mineralogie 12(3-4):279-326.
^ R. E. Fordyce and C. de Muizon. 2001. Evolutionary history of the cetaceans: a review. Secondary Adaptations of Tetrapods to Life in the Water 169-233.
^ O. Lambert. 2005. Systematics and phylogeny of the fossil beaked whales Ziphirostrum du Bus, 1868 and Choneziphius Duvernoy, 1851 (Mammalia, Cetacea, Odontoceti), from the Neogene of Antwerp (North of Belgium). Geodiversitas 27(3):443-497.
^ J. H. Geisler, M. R. McGowen, G. Yang and J. Gatesy. 2011. A supermatrix analysis of genomic, morphological, and paleontological data from crown Cetacea. BMC Evolutionary Biology 11(112):1-33.
^ Olivier Lambert, Stephen J. Godfrey & Erich M. G. Fitzgerald (2018) Yaquinacetus meadi, a new latest Oligocene–early Miocene dolphin (Cetacea, Odontoceti, Squaloziphiidae, fam. nov.) from the Nye Mudstone (Oregon, U.S.A.), Journal of Vertebrate Paleontology, 38:6, DOI: 10.1080/02724634.2018.1559174

Odontocete genera

Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Artiodactyla
Infraorder: Cetacea

Odontoceti

†Agorophius †Ankylorhiza †Archaeodelphis †Argyrocetus †Ashleycetus †Caolodelphis †Chilcacetus †Enigmatocetus †Eoplatanista †Inticetus †Macrodelphinus †Microcetus †Mirocetus †Neosqualodon †Olympicetus †Papahu †Patriocetus †Phoberodon †Phococetus †Prosqualodon †Sachalinocetus †Simocetus †Sulakocetus †Tangaroasaurus
†Xenorophidae	Albertocetus Cotylocara Echovenator Inermorostrum Xenorophus
†Waipatiidae	Awamokoa Ediscetus Nihohae Nihoroa Otekaikea Urkudelphis Waipatia
†Squalodontidae	Eosqualodon Squalodon
†Squaloziphiidae	Crisocetus? Squaloziphius Yaquinacetus
†Eurhinodelphinidae	Ceterhinops Eurhinodelphis Iniopsis Mycteriacetus Phocaenopsis Schizodelphis Vanbreenia Xiphiacetus Ziphiodelphis
Crown-Odontoceti	see below↓

Crown-Odontoceti

Physeteroidea

†Acrophyseter †Albicetus †Brygmophyseter †Eudelphis †Hoplocetus †Livyatan †Miophyseter †Rhaphicetus †Scaldicetus †Zygophyseter
Kogiidae	†Aprixokogia Kogia †Koristocetus †Nanokogia †Platyscaphokogia †Pliokogia †Praekogia †Scaphokogia †Thalassocetus?
Physeteridae	†Aulophyseter †Cozzuoliphyseter †Diaphorocetus †Idiophyseter †Idiorophus †Orycterocetus Physeter †Physeterula †Placoziphius †Thalassocetus?

Platanistoidea

†Aondelphis †Dolgopolis †Ensidelphis †Perditicetus †Eurhinodelphinidae? †Squalodontidae? †Waipatiidae?
†Allodelphinidae?	Allodelphis Arktocara Goedertius Ninjadelphis Zarhinocetus
†Squalodelphinidae?	Furcacetus Huaridelphis Macrosqualodelphis Medocinia Notocetus Phocageneus Squalodelphis
Platanistidae	†Araeodelphis †Dilophodelphis †Grimadelphis †Pachyacanthus? Platanista †Pomatodelphis †Prepomatodelphis †Zarhachis?

Ziphiidae

†Aporotus †Beneziphius †Caviziphius †Chavinziphius †Chimuziphius †Choneziphius †Dagonodum †Globicetus †Imocetus †Messapicetus †Nazcacetus †Nenga †Ninoziphius †Notoziphius †Pterocetus Tasmacetus †Tusciziphius †Xhosacetus †Ziphirostrum
Berardiinae	†Archaeoziphius Berardius †Microberardius
Ziphiinae	†Izikoziphius Ziphius
Hyperoodontinae	†Africanacetus Hyperoodon †Ihlengesi Indopacetus Mesoplodon

Delphinida	see below↓

Delphinida

†Albireo †Atocetus †Brevirostrodelphis †Cammackacetus †Delphinodon †Hadrodelphis †Herbeinodelphis †Liolithax †Lophocetus †Macrokentriodon †Miminiacetus †Odobenocetops †Pictodelphis †Pithanodelphis †Tagicetus †Westmorelandelphis
†Kentriodontidae	Belonodelphis? Kampholophos Kentriodon Platysvercus Sophianacetus Rudicetus Wimahl
Lipotidae	Lipotes †Parapontoporia †Prolipotes

Inioidea

†Awadelphis
Iniidae	†Brujadelphis †Goniodelphis Inia †Ischyrorhynchus †Isoninia? †Isthminia †Kwanzacetus †Meherrinia †Plicodontinia †Saurocetes
Pontoporiidae	†Atocetus? †Auroracetus †Brachydelphis †Piscorhynchus †Pliopontos †Pontistes †Protophocaena †Samaydelphis †Scaldiporia †Stenasodelphis Pontoporia

Delphinoidea

Monodontidae	†Bohaskaia †Casatia Delphinapterus †Denebola †Haborodelphis Monodon
Phocoenidae	†Archaeophocaena †Australithax †Haborophocoena †Lomacetus †Miophocaena Neophocaena †Numataphocoena Phocoena Phocoenoides †Piscolithax †Pterophocaena? †Semirostrum †Septemtriocetus

Delphinidae

†Arimidelphis †Australodelphis †Eodelphinus †Etruridelphis †Hemisyntrachelus Lagenorhynchus †Norisdelphis Orcinus †Platalearostrum †Pliodelphis †Pterophocaena?
Lissodelphininae	Cephalorhynchus Lissodelphis
Delphininae	Delphinus Lagenodelphis Sotalia Sousa Stenella Tursiops
Globicephalinae	Feresa Globicephala Grampus Orcaella Peponocephala Pseudorca Steno