Tolecusatellitidae
Virus classification
(unranked):	Virus
Family:	*Tolecusatellitidae*
Genera
Betasatellite – 119 species Deltasatellite – 12 species

Tolecusatellitidae is a incertae sedis ssDNA/ssDNA(+) family of biological satellites. The family contains two genera and 131 species.^[1] This family of viruses depend on the presence of another virus (helper viruses) to replicate their genomes, as such they have minimal genomes with very low genomic redundancy. ^[2]

Name

The name Tolecusatellitidae is a combination of Tolecu, from the first DNA satellite shown to be associated with Tomato leaf curl virus and satellite, the fact that it is a satellite.^[3]

Taxonomy

The species are ssDNA unless specified ssDNA(+). There are 62 ssDNA(+) and 69 ssDNA satellites.

Betasatellite

Ageratum leaf curl Buea betasatellite
Ageratum leaf curl Cameroon betasatellite
Ageratum yellow leaf curl betasatellite
Ageratum yellow vein betasatellite
Ageratum yellow vein China betasatellite ssDNA(+)
Ageratum yellow vein Sri Lanka betasatellite
Alternanthera yellow vein betasatellite
Andrographis yellow vein leaf curl betasatellite
Bhendi yellow vein mosaic betasatellite
Cardiospermum yellow leaf curl betasatellite
Chili leaf curl betasatellite
Chili leaf curl Jaunpur betasatellite
Chili leaf curl Sri Lanka betasatellite
Codiaeum leaf curl betasatellite ssDNA(+)
Cotton leaf curl Bahraich betasatellite ssDNA(+)
Cotton leaf curl Bangalore betasatellite 1 ssDNA(+)
Cotton leaf curl Bangalore betasatellite 2 ssDNA(+)
Cotton leaf curl Bangalore betasatellite 3 ssDNA(+)
Cotton leaf curl Bangalore betasatellite 4 ssDNA(+)
Cotton leaf curl Burkina Faso betasatellite ssDNA(+)
Cotton leaf curl Gezira betasatellite
Cotton leaf curl Kashmir betasatellite ssDNA(+)
Cotton leaf curl Multan betasatellite
Cotton leaf curl Tandojam betasatellite ssDNA(+)
Croton yellow vein mosaic betasatellite
Emilia yellow vein betasatellite ssDNA(+)
Emilia yellow vein Fujian betasatellite ssDNA(+)
Erectites yellow mosaic betasatellite ssDNA(+)
Eupatorium yellow vein betasatellite
Eupatorium yellow vein mosaic betasatellite
French bean leaf curl betasatellite
Hedyotis yellow mosaic betasatellite
Hibiscus vein enation betasatellite ssDNA(+)
Honeysuckle yellow vein betasatellite
Honeysuckle yellow vein mosaic betasatellite
Honeysuckle yellow vein mosaic Ibaraki betasatellite ssDNA(+)
Honeysuckle yellow vein mosaic Nara betasatellite ssDNA(+)
Kenaf leaf curl betasatellite ssDNA(+)
Leucas zeylanica yellow vein betasatellite ssDNA(+)
Lindernia anagallis yellow vein betasatellite ssDNA(+)
Ludwigia leaf distortion betasatellite 1 ssDNA(+)
Ludwigia leaf distortion betasatellite 2 ssDNA(+)
Ludwigia leaf distortion betasatellite 3 ssDNA(+)
Ludwigia yellow vein betasatellite ssDNA(+)
Malvastrum leaf curl betasatellite ssDNA(+)
Malvastrum yellow vein betasatellite ssDNA(+)
Malvastrum yellow vein Cambodia betasatellite ssDNA(+)
Malvastrum yellow vein Yunnan betasatellite 1 ssDNA(+)
Malvastrum yellow vein Yunnan betasatellite 2 ssDNA(+)
Mirabilis leaf curl betasatellite
Momordica yellow mosaic betasatellite
Mungbean yellow mosaic betasatellite
Okra leaf curl betasatellite ssDNA(+)
Okra leaf curl Oman betasatellite
Papaya leaf curl betasatellite
Papaya leaf curl China betasatellite
Papaya leaf curl Gandhinagar betasatellite ssDNA(+)
Papaya leaf curl India betasatellite
Papaya leaf curl India betasatellite 2 ssDNA(+)
Pea leaf distortion betasatellite ssDNA(+)
Radish leaf curl betasatellite ssDNA(+)
Rhynchosia yellow mosaic betasatellite
Rose leaf curl betasatellite
Sida leaf curl betasatellite ssDNA(+)
Sida yellow mosaic betasatellite ssDNA(+)
Sida yellow vein Barrackpore betasatellite ssDNA(+)
Sida yellow vein Madurai betasatellite ssDNA(+)
Sida yellow vein Vietnam betasatellite 1 ssDNA(+)
Sida yellow vein Vietnam betasatellite 2 ssDNA(+)
Siegesbeckia yellow vein betasatellite
Siegesbeckia yellow vein betasatellite 2 ssDNA(+)
Siegesbeckia yellow vein Guangxi betasatellite ssDNA(+)
Tobacco curly shoot betasatellite
Tobacco leaf chlorosis betasatellite ssDNA(+)
Tobacco leaf curl betasatellite
Tobacco leaf curl Japan betasatellite
Tobacco leaf curl Patna betasatellite
Tobacco leaf curl Sheikhupura betasatellite ssDNA(+)
Tobacco leaf curl Yunnan betasatellite ssDNA(+)
Tomato leaf curl Bangalore betasatellite
Tomato leaf curl Bangalore betasatellite 2 ssDNA(+)
Tomato leaf curl Bangladesh betasatellite
Tomato leaf curl betasatellite ssDNA(+)
Tomato leaf curl betasatellite 2 ssDNA(+)
Tomato leaf curl Bundi betasatellite
Tomato leaf curl China betasatellite
Tomato leaf curl Gandhinagar betasatellite
Tomato leaf curl Ghana betasatellite 1 ssDNA(+)
Tomato leaf curl Ghana betasatellite 2 ssDNA(+)
Tomato leaf curl Hajipur betasatellite ssDNA(+)
Tomato leaf curl India betasatellite ssDNA(+)
Tomato leaf curl Java betasatellite
Tomato leaf curl Joydebpur betasatellite
Tomato leaf curl Joydebpur betasatellite 2 ssDNA(+)
Tomato leaf curl Karnataka betasatellite ssDNA(+)
Tomato leaf curl Laguna betasatellite
Tomato leaf curl Laos betasatellite
Tomato leaf curl Lucknow betasatellite ssDNA(+)
Tomato leaf curl Malaysia betasatellite
Tomato leaf curl Nepal betasatellite
Tomato leaf curl Pakistan betasatellite ssDNA(+)
Tomato leaf curl Panipat betasatellite ssDNA(+)
Tomato leaf curl Patna betasatellite
Tomato leaf curl Philippine betasatellite
Tomato leaf curl Pune betasatellite ssDNA(+)
Tomato leaf curl Ranchi betasatellite ssDNA(+)
Tomato leaf curl Sri Lanka betasatellite
Tomato leaf curl Togo betasatellite ssDNA(+)
Tomato leaf curl Yemen betasatellite
Tomato yellow dwarf betasatellite ssDNA(+)
Tomato yellow leaf curl China betasatellite
Tomato yellow leaf curl Shandong betasatellite
Tomato yellow leaf curl Thailand betasatellite
Tomato yellow leaf curl Vietnam betasatellite
Tomato yellow leaf curl Yunnan betasatellite
Vernonia crinkle betasatellite ssDNA(+)
Vernonia yellow vein betasatellite
Vernonia yellow vein Fujian betasatellite
Zinnia leaf curl betasatellite ssDNA(+)

Deltasatellite

Croton yellow vein deltasatellite
Desmodium leaf distortion deltasatellite ssDNA(+)
Malvastrum leaf curl deltasatellite
Sida golden yellow vein deltasatellite 1
Sida golden yellow vein deltasatellite 2
Sida golden yellow vein deltasatellite 3
sweet potato leaf curl deltasatellite 1
sweet potato leaf curl deltasatellite 2
sweet potato leaf curl deltasatellite 3
tomato leaf curl deltasatellite
tomato yellow leaf distortion deltasatellite 1
tomato yellow leaf distortion deltasatellite 2

References

^ "Virus Taxonomy: 2022 Release". International Committee on Taxonomy of Viruses (ICTV). March 2023. Retrieved 19 August 2023.
^ Silva JM, Pratas D, Caetano T, Matos S (August 2022). "The complexity landscape of viral genomes". GigaScience. 11: giac079. doi:10.1093/gigascience/giac079. PMC 9366995. PMID 35950839.
^ "Proposal 2016.021a-kP.A.v2.Tolecusatellitidae.pdf PDF" (PDF). International Committee on Taxonomy of Viruses (ICTV). Retrieved 19 August 2023.

Life, non-cellular life, and comparable structures

Cellular life

"Prokaryota"	Archaea Bacteria Mitochondrion Plastid LUCA
Eukaryota	Animalia Fungi Plantae 'Protista'
Incertae sedis	Parakaryon Microbes with highly unusual DNA (?)

Non-cellular life

Virus

Viroid

Satellite

Realms

Unassigned

Classes	Naldaviricetes
Families	Alphasatellitidae Ampullaviridae Anelloviridae Avsunviroidae Bicaudaviridae Clavaviridae Finnlakeviridae Fuselloviridae Globuloviridae Guttaviridae Halspiviridae Ovaliviridae Plasmaviridae Polydnaviridae Portogloboviridae Pospiviroidae Spiraviridae Thaspiviridae Tolecusatellitidae
Genera	Dinodnavirus Rhizidiovirus

Other

Nanobe (?)
Obelisk

Comparable
structures

Bio-like structure
Cancer cell
- HeLa
- Clonally transmissible cancer
Cosmid
Defective interfering particle
Endogenous viral element
Fosmid
Integrative and conjugative element
Jeewanu
"Nanobacterium"
Phagemid
Plasmid
Prion
- Fungal prion
Proteinoid microsphere
Retroelements not elsewhere classified
- Retron
- Diversity-generating retroelement
- Telomerase reverse transcriptase
- Reverse transcriptase-related cellular gene
Ribozyme
Spiegelman's Monster
Tandem repeat
Transposable element
- Retroposon
Transpoviron
Xenobot