Trionyx Temporal range: Miocene to recent, 23–0 Ma^[1] PreꞒ Ꞓ O S D C P T J K Pg N

T. triunguis, the only extant species
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Cryptodira
Family:	Trionychidae
Subfamily:	Trionychinae
Genus:	Trionyx Geoffroy Saint-Hilaire, 1809
Synonyms
Aspidonectes Wagler, 1830

Trionyx is a genus of softshell turtles belonging to the family Trionychidae. In the past many species in the family were classified in this genus, but today T. triunguis, the African or Nile softshell turtle, is the only extant softshell still classified as Trionyx. The other species still assigned to this genus are only known from fossils. T. triunguis is a relatively large, aquatic piscivore.

Species

The list of species follows a 2017 review of the Trionychidae by Georgios L. Georgalis and Walter G. Joyce.^[2]

Trionyx triunguis (Forsskål, 1775) – African or Nile softshell turtle
† Trionyx pliocenicus Fucini, 1912 – Pliocene of Tuscany (Italy)^[2]
† Trionyx vindobonensis (Peters, 1855) – Miocene of Vienna (Austria)^[2]

The following species are considered valid trionychid taxa, but with uncertain phylogenetic relationships. They are only referred to Trionyx provisionally due to its historic status as a wastebasket taxon.^[2] The species from North America follow a 2015 review by Natasha S. Vitek and Walter G. Joyce^[3] and the species from Europe and Asia follow the 2017 review by Georgalis and Joyce.^[2]

† "Trionyx" admirabilis (Hay, 1905) – Paleocene of New Mexico (United States)^[3]
† "Trionyx" aequa (Hay, 1908) – Eocene of Wyoming (United States)^[3]
† "Trionyx" allani (Gilmore, 1923) – Late Cretaceous of Alberta (Canada)^[3]
† "Trionyx" austerus (Hay, 1908) – Late Cretaceous of New Mexico (United States)^[3]
† "Trionyx" baynshirensis Danilov et al., 2014 – Middle Cretaceous of Dornogovi Aimag (Mongolia)^[2]
† "Trionyx" beecheri (Hay, 1904) – Late Cretaceous of Wyoming (United States)^[3]
† "Trionyx" boulengeri Reinach, 1900 – Oligocene of Rhineland-Palatinate (Germany)^[2]
† "Trionyx" capellinii Negri, 1892 – Eocene of Veneto (Italy)^[2]
† "Trionyx" dissolutus Vitek & Danilov, 2014 – Late Cretaceous of Navoiy Region (Uzbekistan)^[2]
† "Trionyx" ellipticus (Hay, 1908) – Eocene of Utah (United States)^[3]
† "Trionyx" eloisae (Gilmore, 1919) – Paleocene of New Mexico (United States)^[3]
† "Trionyx" foveatus Leidy, 1857 – Late Cretaceous of Montana (United States)^[3]
† "Trionyx" gilbentuensis Danilov et al., 2014 – Late Cretaceous of Ömnögovi Aimag (Mongolia)^[2]
† "Trionyx" gobiensis Danilov et al., 2014 – Late Cretaceous of Ömnögovi Aimag (Mongolia)^[2]
† "Trionyx" gregarius (Gilmore, 1934) – Eocene of Inner Mongolia (China)^[2]
† "Trionyx" henrici Owen, 1849 – Eocene of Hampshire (United Kingdom)^[2]
† "Trionyx" ikoviensis Danilov et al., 2011 – Eocene of Luhansk (Ukraine)^[2]
† "Trionyx" impressus (Yeh, 1963) – Eocene of Yuangdong Province (China)^[2]
† "Trionyx" latus (Gilmore, 1919) – Late Cretaceous of Alberta (Canada)^[3]
† "Trionyx" leucopotamicus Cope, 1891 – Eocene of Saskatchewan (Canada)^[3]
† "Trionyx" johnsoni Gilmore, 1931 – Eocene of Inner Mongolia (China)^[2]
† "Trionyx" kansaiensis Vitek & Danilov, 2010 – Late Cretaceous of Khodzhent Province (Tajikistan)^[2]
† "Trionyx" linchuensis (Yeh, 1962) – Eocene of Shandong Province (China)^[2]
† "Trionyx" messelianus Reinach, 1900 – Eocene of Hesse (Germany)^[2]
† "Trionyx" miensis Okazaki and Yoshida, 1977 – Pliocene of Mie Prefecture (Japan)^[2]
† "Trionyx" miocaenus (Matthew, 1924) – Eocene of Wyoming (United States)^[3]
† "Trionyx" minusculus (Chkhikvadze, 1973) – Eocene of East Kazakhstan Region (Kazakhstan)^[2]
† "Trionyx" mira (Hay, 1908) – Eocene of Wyoming (United States)^[3]
† "Trionyx" ninae Chkhikvadze, 1971 – Oligocene of Karagandy Region (Kazakhstan)^[2]
† "Trionyx" onomatoplokos Georgalis & Joyce, 2017 – Late Cretaceous of Kyzylorda Region (Kazakhstan)^[2]
† "Trionyx" reesidei (Gilmore, 1919) – Paleocene of New Mexico (United States)^[3]
† "Trionyx" riabinini Kuznetsov & Chkhikvadze, 1987 – Late Cretaceous of Kyzylorda Region (Kazakhstan)^[2]
† "Trionyx" robustus (Gilmore, 1919) – Late Cretaceous of New Mexico (United States)^[3]
† "Trionyx" shiluutulensis Danilov et al., 2014 – Late Cretaceous of Ömnögovi Aimag (Mongolia)^[2]
† "Trionyx" silvestris Walker & Moody, 1974 – Eocene of Kent (United Kingdom)^[2]
† "Trionyx" singularis (Hay, 1907) – Paleocene of New Mexico (United States)^[3]
† "Trionyx" uintaensis Leidy, 1873 – Eocene of Wyoming (United States)^[3]
† "Trionyx" vegetus (Gilmore, 1919) – Paleocene of New Mexico (United States)^[3]
† "Trionyx" yixiensis Li, Tong et al., 2015 – Early Cretaceous of Heilongjiang Province (China)^[2]

† = extinct

References

External links

Trionychidae family

Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Subclass: Anapsida
Order: Testudines
Superfamily: Trionychia

Genera

Species of the Trionychidae family

Amyda

Asiatic softshell turtle
†Amyda gregaria
Amyda ornata

Apalone

†Axestemys

†Axestemys byssina
†Axestemys cerevisia
†Axestemys montinsana
†Axestemys quinni
†Axestemys splendida
†Axestemys vittata

Chitra

Cyclanorbis

Cycloderma

Dogania

Malayan softshell turtle

†Gilmoremys

†Gilmoremys gettyspherensis
†Gilmoremys lancensis

†Hutchemys

†Hutchemys arctochelys
†Hutchemys rememdium
†Hutchemys tetanetron
†Hutchemys sterea
†Hutchemys acupictus

†Khunnuchelys

†Khunnuchelys erinhotensis
†Khunnuchelys kizylkumensis
†Khunnuchelys lophorhothon

Lissemys

Nilssonia

†Palaeoamyda

†Palaeoamyda messeliana

Palea

Wattle-necked softshell turtle

Pelochelys

Pelodiscus

Rafetus

Trionyx

Phylogenetic arrangement of turtles based on turtles of the world 2017 update: Annotated checklist and atlas of taxonomy, synonymy, distribution, and conservation status. †=extinct.

Testudines

Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Diapsida
Order: Testudines

Genus

Chelonioidea
(Sea turtles)

Cheloniidae	†Allopleuron Caretta †Carolinochelys Chelonia †Eochelone Eretmochelys †Gigantatypus †Glarichelys †Itilochelys Lepidochelys †Mexichelys †Miocaretta Natator †Pacifichelys †Syllomus †Tasbacka
Dermochelyidae	†Arabemys †Corsochelys †Cosmochelys Dermochelys †Eosphargis †Mesodermochelys †Psephophorus
	†Euclastes †Peritresius †Procolpochelys †Protosphargis †Puppigerus

Kinosternoidea

Dermatemydidae	Dermatemys
Kinosternidae	Claudius †Hoplochelys Kinosternon Staurotypus Sternotherus

Testudinoidea

Emydidae	†Acherontemys Chrysemys Clemmys Deirochelys Emys Actinemys Emydoidea Glyptemys Graptemys Malaclemys Pseudemys Terrapene Trachemys †Wilburemys
Geoemydidae	Batagur †Banhxeochelys Cuora Cyclemys Geoclemys Geoemyda Hardella Heosemys Leucocephalon Malayemys Mauremys Melanochelys Morenia Notochelys Orlitia Pangshura Rhinoclemmys Sacalia Siebenrockiella Vijayachelys
Platysternidae	Platysternon
Testudinidae	Aldabrachelys Astrochelys Centrochelys Chelonoidis Chersina Cylindraspis †Cymatholcus †Floridemys Geochelone Gopherus †Hadrianus †Hesperotestudo Homopus Indotestudo Kinixys Malacochersus Manouria †Megalochelys †Oligopherus Psammobates Pyxis †Solitudo Stigmochelys †Stylemys Testudo

Trionychia

Carettochelyidae	†Allaeochelys †Anosteira Carettochelys
Trionychidae	Amyda Apalone †Axestemys Chitra Cyclanorbis Cycloderma †Drazinderetes Dogania †Gilmoremys †Hutchemys †Khunnuchelys Lissemys Nilssonia Palea †Palaeoamyda Pelochelys Pelodiscus Rafetus Trionyx
	†Basilochelys †Sinaspideretes

Chelydridae	Chelydra †Chelydrops †Chelydropsis †Emarginachelys †Macrocephalochelys Macrochelys †Planiplastron †Protochelydra
†Nanhsiungchelyidae	†Anomalochelys †Basilemys †Jiangxichelys
†Protostegidae	†Alienochelys †Archelon †Atlantochelys †Bouliachelys †Calcarichelys †Cratochelone †Desmatochelys †Iserosaurus †Notochelone †Ocepechelon †Pneumatoarthrus †Protostega †Rhinochelys †Santanachelys †Terlinguachelys
	†Adocus †Argillochelys †Bashuchelys †Ctenochelys †Prionochelys †Toxochelys

Pleurodira

†Araripemydidae	†Araripemys
†Bothremydidae	†Araiochelys †Arenila †Azabbaremys †Bothremys †Cearachelys †Chedighaii †Chupacabrachelys †Eotaphrosphys †Foxemys †Galianemys †Ilatardia †Inaechelys †Itapecuruemys †Jainemys †Kinkonychelys †Kurmademys †Labrostochelys †Nigeremys †Phosphatochelys †Polysternon †Puentemys †Rosasia †Rhothonemys †Sankuchemys †Taphrosphys †Ummulisani †Zolhafah
Chelidae	Acanthochelys Chelodina Chelus Elseya Elusor Emydura Hydromedusa †Lomalatachelys Mesoclemmys Myuchelys Phrynops Platemys †Prochelidella Pseudemydura Rheodytes Rhinemys †Yaminuechelys
Pelomedusidae	Pelomedusa Pelusios
Podocnemididae	†Albertwoodemys †Bauruemys †Brontochelys †Caninemys †Carbonemys †Cerrejonemys †Cordichelys Erymnochelys †Lapparentemys †Latentemys Peltocephalus Podocnemis †Stupendemys
†Sahonachelyidae	†Sahonachelys †Sokatra

	†Caribemys †Caririemys †Tacuarembemys

Phylogenetic arrangement of turtles based on Turtles of the World 2017 Update: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status. † = extinct.

See also List of Testudines families

Taxon identifiers

Taxon identifiers
Trionyx	Wikidata: Q5079121 Wikispecies: Trionyx BOLD: 247016 CoL: 64853 EoL: 46559485 GBIF: 2442547 iNaturalist: 39552 ITIS: 173846 NBN: NHMSYS0000375799 NCBI: 13734 Open Tree of Life: 143397 Paleobiology Database: 37682 WoRMS: 397353