Family of gastropods
Urocyclidae is a family of air-breathing land snails, semi-slugs and land slugs, terrestrial pulmonate gastropod mollusks in the superfamily Helicarionoidea, (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).[2]
Some species within this family make and use love darts before mating.[citation needed]
Distribution
The Urocyclidae are distributed in the Afrotropical realm, including Madagascar, the Comores and the islands of the Gulf of Guinea.[3][4]
Taxonomy
The family Urocyclidae consists of 3 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005),[2] that follows classification of Urocyclinae by Van Goethem (1977):[5]
- subfamily Rhysotininae Schileyko, 2002
- subfamily Sheldoniinae Connolly, 1925 (1912)[6] - synonyms: Peltatinae Godwin-Austen, 1912;[7] Trochonanininae Connolly, 1912;[8] Trochozonitinae Iredale, 1914;[9] Ledoulxiinae Pilsbry, 1919;[10] Gymnarionidae Von Mol, 1970;[11] Rhysotinidae Schileyko, 2002; Zonitarionini Schileyko, 2002; Acantharionini Schileyko, 2002
- subfamily Urocyclinae Simroth, 1889
- tribe Urocyclini Simroth, 1889 - synonyms: Atoxonini Schileyko, 2002; Buettneriini Schileyko, 2002
- tribe Dendrolimacini Van Goethem, 1977[5]
- tribe Leptichnini Van Goethem, 1977[5]
- tribe Upembellini Van Goethem, 1977[5]
Genera
Genera within the family Urocyclidae include:
- unsorted
- † Koruella Pickford, 2009
- Subfamily Rhysotininae Schileyko, 2002
- Rhysotina Ancey, 1887[2]
- Thomeonanina Germain, 1909 : synonym of Rhysotina Ancey, 1887 (objective synonym)
- Subfamily Sheldoniinae
- Acantharion Binder & Tillier, 1985[2]
- Africarion Godwin-Austen, 1883
- Amatarion Van Mol, 1970
- Angustivestis Pilsbry, 1919
- Aspidelus Morelet, 1884
- Aspidotomium Degner, 1932
- Belonarion Pilsbry, 1919
- Bloyetia Bourguignat, 1890
- Camerunarion Van Mol, 1970
- Carinazingis van Bruggen & de Winter, 1990
- Centrafricarion Van Mol, 1970 [11]
- Chlamydarion Van Mol, 1968
- Colparion Laidlaw, 1938
- Degneria Verdcourt, 1956
- Entagaricus Pilsbry, 1919
- Estria Poirier, 1887
- Falloonella Preston, 1914
- Fuellebornia Verdcourt, 1998
- Granularion Germain, 1912
- Gymnarion Pilsbry, 1919[2]
- Hamya Bourguignat, 1885 †
- Kerkophorus Godwin-Austen, 1912
- Lacrimarion Connolly, 1939
- Malagarion Tillier, 1979
- Mesafricarion Pilsbry, 1919
- Microkerkus Godwin-Austen, 1912
- Montanobloyetia Verdcourt, 1961
- Morrumbalia Verdcourt, 1998
- Paragranularion Van Mol, 1970
- Percivalia Preston, 1914
- Plicatonanina Verdcourt, 1961
- Principicochlea D. Holyoak & G. Holyoak, 2020
- Principitrochoidea D. Holyoak & G. Holyoak, 2020
- Ptilototheca Herbert, 2016
- Rhopalogonium Degner, 1932
- Selatodryas Herbert, 2017
- Senegalarion Van Mol, 1970
- Sheldonia Ancey, 1887 - the type genus of the subfamily Sheldoniinae[2][12]
- Sjostedtina Verdcourt, 1961
- Sylvarion Van Mol, 1970
- Thapsia Albers, 1860
- Thielarion Van Mol, 1970
- Thomithapsia D. Holyoak & G. Holyoak, 2020
- Thomitrochoidea D. Holyoak & G. Holyoak, 2020
- Tresia Van Goethem, 1975
- Trochonanina Mousson, 1869[2]
- Trochozonites Pfeffer, 1883[2]
- Tropidocochlion Verdcourt, 1998
- Verdcourtia Van Mol, 1970
- Verrucarion Van Mol, 1970 [13]
- Zingis E. von Martens, 1878
- Zonitarion Pfeffer, 1883[2]
- Ledoulxia Bourguignat, 1885:[2] synonym of Trochonanina Mousson, 1869
- Peltatus Godwin-Austen, 1908:[2] synonym of Sheldonia Ancey, 1887
- Subfamily Urocyclinae
Tribe Dendrolimacini
- Dendrolimax Heynemann, 1868 - the type genus of the tribe Dendrolimacini[2]
Tribe Leptichnini
- Leptichnus Simroth, 1896 - the type genus of the tribe Leptichnini[2]
Tribe Upembellini
- Leptichnoides Van Goethem, 1975
- Upembella Van Goethem, 1969 - the type genus of the tribe Upembellini[2]
tribe Urocyclini
- Anisotoxon Van Goethem, 1975
- Atoxon Simroth, 1888[2]
- Atoxonoides Van Goethem, 1973
- Atrichotoxon Simroth, 1910
- Buettneria Simroth, 1910[14]
- Bukobia Simroth, 1896
- Elisolimax Cockerell, 1893
- Emphysetes Verdcourt, 2003
- Microcyclus Simroth, 1896
- Nupnus Van Goethem, 1975
- Pembatoxon Van Goethem, 1975
- Phaneroporus Simroth, 1888
- Polytoxon Simroth, 1897
- Pseudatoxon Van Goethem, 1975
- Tanzalimax Rowson, Paustian & Van Goethem, 2017
- Trichotoxon Simroth, 1888
- Udzungwalimax Rowson, Paustian & Van Goethem, 2017
- Urocyclus Gray, 1864 - the type genus of the family Urocyclidae[2]
