.mw-parser-output .hidden-begin{box-sizing:border-box;width:100%;padding:5px;border:none;font-size:95%}.mw-parser-output .hidden-title{font-weight:bold;line-height:1.6;text-align:left}.mw-parser-output .hidden-content{text-align:left}This article may be expanded with text translated from the corresponding article in Spanish. (December 2009) Click [show] for important translation instructions. View a machine-translated version of the Spanish article. Machine translation, like DeepL or Google Translate, is a useful starting point for translations, but translators must revise errors as necessary and confirm that the translation is accurate, rather than simply copy-pasting machine-translated text into the English Wikipedia. Do not translate text that appears unreliable or low-quality. If possible, verify the text with references provided in the foreign-language article. You must provide copyright attribution in the edit summary accompanying your translation by providing an interlanguage link to the source of your translation. A model attribution edit summary is Content in this edit is translated from the existing Spanish Wikipedia article at [[:es:Gunnerales]]; see its history for attribution. You should also add the template ((Translated|es|Gunnerales)) to the talk page. For more guidance, see Wikipedia:Translation.

Gunnera tinctoria0.jpg
Scientific classification e
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Core eudicots
Order: Gunnerales
Takht. ex Reveal[1]

The Gunnerales are an order of flowering plants. In the APG III system (2009) and APG IV system (2016) it contains two genera: Gunnera (in family Gunneraceae) and Myrothamnus (in family Myrothamnaceae). In the Cronquist system (1981), the Gunneraceae were in the Haloragales and Myrothamnaceae in the Hamamelidales.[1] DNA analysis was definitive, but the grouping of the two families was a surprise, given their very dissimilar morphologies. In Cronquist's old system (1981, 1988), and Takhtajan's (1997), the Gunneraceae were in the Rosidae, and the Myrothamnaceae were in the Hamamelids. In modern classification systems such as APG III and APG IV this order was the first to derive from the core eudicots.[1][2]


Both families contain ellagic acid. Phloem cells contain a large number of plastids and the leaves have dented borders.

The plants are dioecious, have small flowers without perianth, and the stigma is at least weakly secretory. Gunnerales characters shared with the core of the eudicots are cyanogenesis via phenylalanine, metabolic pathways of isoleucine or valine, presence of the DNA sequence of PI-dB motif, 9 and is common to suffer a small deletion in the sequence of 18S ribosomal DNA. The characters which it shares with the core of eudicotyledons, and also with Buxales and Trochodendrales are: absence of benzylisoquinoline alkaloids, euAP3 + TM6 genes (gene duplication paleoAP3: Class B), and loss of the mitochondrial gene rps2.


Despite being related, the Myrothamnaceae and the Gunneraceae look very different:

Both have flowers without perianth, but the details of pollen (e.g. Zavada and Dilcher see 1986 10, Wanntorp et al. 2004th 2004b 11 and 12) differ. In Wilkinson 2000 13 is a table of differences.


  1. ^ a b c Angiosperm Phylogeny Group (2009). "An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG III". Botanical Journal of the Linnean Society. 161 (2): 105–121. doi:10.1111/j.1095-8339.2009.00996.x.
  2. ^ Angiosperm Phylogeny Group (2016). "An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG IV". Botanical Journal of the Linnean Society. 181 (1): 1–20. doi:10.1111/boj.12385. ISSN 0024-4074.((cite journal)): CS1 maint: uses authors parameter (link)