Memetics is a theory of the evolution of culture based on Darwinian principles with the meme as the unit of culture. The term ‘meme’ was coined by biologist Richard Dawkins in his 1976 book The Selfish Gene,[1] to illustrate the principle that he later called ‘Universal Darwinism’. All evolutionary processes depend on information being copied, varied, and selected; a process also known as variation with selective retention. The information that is copied is called the replicator, and genes are the replicator for biological evolution. Dawkins proposed that the same process drives cultural evolution, and he called this second replicator the ‘meme’. He gave as examples, tunes, catchphrases, fashions, and technologies. Like genes, memes are selfish replicators and have causal efficacy, in other words their properties influence their chances of being copied and passed on. Some succeed because they are valuable or useful to their human hosts while others are more like viruses.

Just as genes can work together to form co-adapted gene complexes, so groups of memes acting together form co-adapted meme complexes or memeplexes. Memeplexes include (among many other things) languages, traditions, scientific theories, financial institutions, and religions. Dawkins famously referred to religions as ‘viruses of the mind’.

Among proponents of memetics are psychologist Susan Blackmore, author of The Meme Machine, who argues that when our ancestors began imitating behaviours, they let loose a second replicator and co-evolved to become the 'meme machines' that copy, vary, and select memes in culture.[2] Philosopher Daniel Dennett develops memetics extensively, notably in his books Darwin’s Dangerous Idea,[3] and From bacteria to Bach and Back.[4] He describes the units of memes as “the smallest elements that replicate themselves with reliability and fecundity.”[5] and claims that “Human consciousness is itself a huge complex of memes.”[6] In The beginning of Infinity,[7] physicist David Deutsch contrasts static societies that depend on anti-rational memes suppressing innovation and creativity, with dynamic societies based on rational memes that encourage enlightenment, values, scientific curiosity, and progress.

Criticisms of memetics include claims that memes do not exist, that the analogy with genes is false, that the units cannot be specified, that culture does not evolve through imitation, and that the sources of variation are intelligently designed rather than random. Critics of memetics include biologist Stephen Jay Gould who calls memetics a ‘meaningless metaphor’. Philosopher Dan Sperber argues against memetics as a viable approach to cultural evolution because cultural items are not directly copied or imitated but are re-produced.[8] Anthropologist Robert Boyd and biologist Peter Richerson work within the alternative, and more mainstream, field of cultural evolution theory and gene-culture coevolution.[9] Dual-inheritance theory has much in common with memetics but rejects the idea that memes are replicators. From this perspective, memetics is seen as just one of several approaches to cultural evolution and one that is generally considered less useful than the alternatives of gene-culture coevolution or dual-inheritance theory. The main difference is that dual-inheritance theory ultimately depends on biological advantage to genes, whereas memetics treats memes as a second replicator in its own right. Memetics also extends to the analysis of digital culture and Internet memes.[10]


In his book The Selfish Gene (1976), the evolutionary biologist Richard Dawkins used the term meme to describe a unit of human cultural transmission analogous to the gene, arguing that replication also happens in culture, albeit in a different sense. While cultural evolution itself is a much older topic, with a history that dates back at least as far as Darwin's era, Dawkins (1976) proposed that the meme is a unit of culture residing in the brain and is the mutating replicator in human cultural evolution. After Dawkins, many discussed this unit of culture as evolutionary "information" which replicates with rules analogous to Darwinian selection.[11] A replicator is a pattern that can influence its surroundings – that is, it has causal agency – and can propagate. This proposal resulted in debate among anthropologists, sociologists, biologists, and scientists of other disciplines. Dawkins himself did not provide a sufficient explanation of how the replication of units of information in the brain controls human behaviour and ultimately culture (the principal topic of the book was genetics). Dawkins apparently did not intend to present a comprehensive theory of memetics in The Selfish Gene, but rather coined the term meme in a speculative spirit. Accordingly, different researchers came to define the term "unit of information" in different ways.

The evolutionary model of cultural information transfer is based on the concept that memes—units of information—have an independent existence, are self-replicating, and are subject to selective evolution through environmental forces.[12] Starting from a proposition put forward in the writings of Dawkins, this model has formed the basis of a new area of study, one that looks at the self-replicating units of culture. It has been proposed that just as memes are analogous to genes, memetics is analogous to genetics.

The modern memetics movement dates from the mid-1980s. A January 1983 "Metamagical Themas" column[13] by Douglas Hofstadter, in Scientific American, was influential – as was his 1985 book of the same name. "Memeticist" was coined as analogous to "geneticist" – originally in The Selfish Gene. Later Arel Lucas suggested that the discipline that studies memes and their connections to human and other carriers of them be known as "memetics" by analogy with "genetics".[14] Dawkins' The Selfish Gene has been a factor in attracting the attention of people of disparate intellectual backgrounds. Another stimulus was the publication in 1991 of Consciousness Explained by Tufts University philosopher Daniel Dennett, which incorporated the meme concept into a theory of the mind. In his 1991 essay "Viruses of the Mind", Richard Dawkins used memetics to explain the phenomenon of religious belief and the various characteristics of organised religions. By then, memetics had also become a theme appearing in fiction (e.g. Neal Stephenson's Snow Crash).

The idea of language as a virus had already been introduced by William S. Burroughs as early as 1962 in his fictional book The Ticket That Exploded, and continued in The Electronic Revolution, published in 1970 in The Job.

The foundation of memetics in its full modern incarnation was launched by Douglas Rushkoff's Media Virus: Hidden Agendas in Popular Culture in 1995,[15] and was accelerated with the publication in 1996 of two more books by authors outside the academic mainstream: Virus of the Mind: The New Science of the Meme by former Microsoft executive turned motivational speaker and professional poker-player Richard Brodie, and Thought Contagion: How Belief Spreads Through Society by Aaron Lynch, a mathematician and philosopher who worked for many years as an engineer at Fermilab. Lynch claimed to have conceived his theory totally independently of any contact with academics in the cultural evolutionary sphere, and apparently was not aware of The Selfish Gene until his book was very close to publication.[citation needed]

Around the same time as the publication of the books by Lynch and Brodie the e-journal Journal of Memetics – Evolutionary Models of Information Transmission[16] (published electronically from 1997 to 2005[17]) first appeared. It was first hosted by the Centre for Policy Modelling at Manchester Metropolitan University. The e-journal soon became the central point for publication and debate within the nascent memeticist community. (There had been a short-lived paper-based memetics publication starting in 1990, the Journal of Ideas edited by Elan Moritz.[18]) In 1999, Susan Blackmore, a psychologist at the University of the West of England, published The Meme Machine, which more fully worked out the ideas of Dennett, Lynch, and Brodie and attempted to compare and contrast them with various approaches from the cultural evolutionary mainstream, as well as providing novel (and controversial) memetics-based theories for the evolution of language and the human sense of individual selfhood.


The term meme derives from the Ancient Greek μιμητής (mimētḗs), meaning "imitator, pretender". The similar term mneme was used in 1904, by the German evolutionary biologist Richard Semon, best known for his development of the engram theory of memory, in his work Die mnemischen Empfindungen in ihren Beziehungen zu den Originalempfindungen, translated into English in 1921 as The Mneme.[19] Until Daniel Schacter published Forgotten Ideas, Neglected Pioneers: Richard Semon and the Story of Memory in 2000, Semon's work had little influence, though it was quoted extensively in Erwin Schrödinger’s 1956 Tarner LectureMind and Matter”. Richard Dawkins (1976) apparently coined the word meme independently of Semon, writing this:

"'Mimeme' comes from a suitable Greek root, but I want a monosyllable that sounds a bit like 'gene'. I hope my classicist friends will forgive me if I abbreviate mimeme to meme. If it is any consolation, it could alternatively be thought of as being related to 'memory', or to the French word même."[20]

David Hull (2001) pointed out Dawkins's oversight of Semon's work. Hull suggests this early work as an alternative origin to memetics by which Dawkins's memetic theory and classicist connection to the concept can be negotiated.

"Why not date the beginnings of memetics (or mnemetics) as 1904 or at the very least 1914? If [Semon's] two publications are taken as the beginnings of memetics, then the development of memetics [...] has been around for almost a hundred years without much in the way of conceptual or empirical advance!"[21]

Despite this, Semon's work remains mostly understood as distinct to memetic origins even with the overt similarities accounted for by Hull.

Internalists and externalists

The memetics movement split almost immediately into two. The first group were those who wanted to stick to Dawkins' definition of a meme as "a unit of cultural transmission". Gibron Burchett, a memeticist responsible for helping to research and co-coin the term memetic engineering, along with Leveious Rolando and Larry Lottman, has stated that a meme can be defined, more precisely, as "a unit of cultural information that can be copied, located in the brain". This thinking is more in line with Dawkins' second definition of the meme in his book The Extended Phenotype. The second group wants to redefine memes as observable cultural artifacts and behaviors. However, in contrast to those two positions, the article "Consciousness in meme machines" by Susan Blackmore rejects neither movement.[22]

These two schools became known as the "internalists" and the "externalists." Prominent internalists included both Lynch and Brodie; the most vocal externalists included Derek Gatherer, a geneticist from Liverpool John Moores University, and William Benzon, a writer on cultural evolution and music. The main rationale for externalism was that internal brain entities are not observable, and memetics cannot advance as a science, especially a quantitative science, unless it moves its emphasis onto the directly quantifiable aspects of culture. Internalists countered with various arguments: that brain states will eventually be directly observable with advanced technology, that most cultural anthropologists agree that culture is about beliefs and not artifacts, or that artifacts cannot be replicators in the same sense as mental entities (or DNA) are replicators. The debate became so heated that a 1998 Symposium on Memetics, organised as part of the 15th International Conference on Cybernetics, passed a motion calling for an end to definitional debates. McNamara demonstrated in 2011 that functional connectivity profiling using neuroimaging tools enables the observation of the processing of internal memes, "i-memes", in response to external "e-memes".[23] This was developed further in a paper "Memetics and Neural Models of Conspiracy Theories" by Duch, where a model of memes as a quasi-stable neural associative memory attractor network is proposed, and a formation of Memeplex leading to conspiracy theories illustrated with the simulation of a self-organizing network.[24]

An advanced statement of the internalist school came in 2002 with the publication of The Electric Meme, by Robert Aunger, an anthropologist from the University of Cambridge. Aunger also organised a conference in Cambridge in 1999, at which prominent sociologists and anthropologists were able to give their assessment of the progress made in memetics to that date. This resulted in the publication of Darwinizing Culture: The Status of Memetics as a Science, edited by Aunger and with a foreword by Dennett, in 2001.[25]


In 2005, the Journal of Memetics ceased publication and published a set of articles on the future of memetics. The website states that although "there was to be a relaunch... after several years nothing has happened".[26] Susan Blackmore left the University of the West of England to become a freelance science-writer and now concentrates more on the field of consciousness and cognitive science. Derek Gatherer moved to work as a computer programmer in the pharmaceutical industry, although he still occasionally publishes on memetics-related matters. Richard Brodie is now climbing the world professional poker rankings. Aaron Lynch disowned the memetics community and the words "meme" and "memetics" (without disowning the ideas in his book), adopting the self-description "thought contagionist". He died in 2005.

Susan Blackmore (2002) re-stated the definition of meme as: whatever is copied from one person to another person, whether habits, skills, songs, stories, or any other kind of information. Further she said that memes, like genes, are replicators in the sense as defined by Dawkins.[27] That is, they are information that is copied. Memes are copied by imitation, teaching and other methods. The copies are not perfect: memes are copied with variation; moreover, they compete for space in our memories and for the chance to be copied again. Only some of the variants can survive. The combination of these three elements (copies; variation; competition for survival) forms precisely the condition for Darwinian evolution, and so memes (and hence human cultures) evolve. Large groups of memes that are copied and passed on together are called co-adapted meme complexes, or memeplexes. In Blackmore's definition, the way that a meme replicates is through imitation. This requires brain capacity to generally imitate a model or selectively imitate the model. Since the process of social learning varies from one person to another, the imitation process cannot be said to be completely imitated. The sameness of an idea may be expressed with different memes supporting it. This is to say that the mutation rate in memetic evolution is extremely high, and mutations are even possible within each and every iteration of the imitation process. It becomes very interesting when we see that a social system composed of a complex network of microinteractions exists, but at the macro level an order emerges to create culture.[citation needed]

Many researchers of cultural evolution regard memetic theory of this time a failed paradigm superseded by dual inheritance theory.[28] Others instead suggest it is not superseded but rather holds a small but distinct intellectual space in cultural evolutionary theory.[29]

"Internet Memetics"

See also: Internet meme

A new framework of Internet Memetics initially borrowed Blackmore's conceptual developments but is effectively a data-driven approach, focusing on digital artifacts. This was led primarily by conceptual developments Colin Lankshear and Michele Knobel (2006) [30] and Limor Shifman and Mike Thelwall (2009).[31] Shiman, in particular, followed Susan Blackmore in rejecting the internalist and externalist debate, however did not offer a clear connection to prior evolutionary frameworks. Later in 2014, she rejected the historical relevance of "information" to memetics. Instead of memes being units of cultural information, she argued information is exclusively delegated to be "the ways in which addressers position themselves in relation to [a meme instance's] text, its linguistic codes, the addressees, and other potential speakers."[32] This is what she called stance, which is analytically distinguished from the content and form of her meme. As such, Shifman's developments can be seen as critical to Dawkins's meme, but also as a somewhat distinct conceptualization of the meme as a communicative system dependent on the internet and social media platforms. By introducing memetics as an internet study there has been a rise in empirical research. That is, memetics in this conceptualization has been notably testable by the application of social science methodologies. It has been popular enough that following Lankshear and Knobel's (2019) review of empirical trends, they warn those interested in memetics that theoretical development should not be ignored, concluding that,

"[R]ight now would be a good time for anyone seriously interested in memes to revisit Dawkins’ work in light of how internet memes have evolved over the past three decades and reflect on what most merits careful and conscientious research attention."[33]

As Lankshear and Knobel show, the Internet Memetic reconceptualization is limited in addressing long-standing memetic theory concerns. It is not clear that existing Internet Memetic theory's departure from conceptual dichotomies between internalist and externalist debate are compatible with most earlier concerns of memetics. Internet Memetics might be understood as a study without an agreed upon theory, as present research tends to focus on empirical developments answering theories of other areas of cultural research. It exists more as a set of distributed studies than a methodology, theory, field, or discipline, with a few exceptions such as Shifman and those closely following her motivating framework.


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Critics contend that some of the proponents' assertions are "untested, unsupported or incorrect."[12] Most of the history of memetic criticism has been directed at Dawkins' earlier theory of memetics framed in The Selfish Gene. There have been some serious criticisms of memetics. Namely, there are a few key points on which most criticisms focus: mentalism, cultural determinism, Darwinian reduction, a lack of academic novelty, and a lack of empirical evidence of memetic mechanisms.

Luis Benitez-Bribiesca points to the lack of memetic mechanisms. He refers to the lack of a code script for memes which would suggest a genuine analogy to DNA in genes. He also suggests the meme mutation mechanism is too unstable which would render the evolutionary process chaotic. That is to say that the "unit of information" which traverses across minds is perhaps too flexible in meaning to be a realistic unit.[34] As such, he calls memetics "a pseudoscientific dogma" and "a dangerous idea that poses a threat to the serious study of consciousness and cultural evolution" among other things.

Another criticism points to memetic triviality. That is, some have argued memetics is derivative of more rich areas of study. One of these cases comes from Peircian semiotics, (e.g., Deacon,[35] Kull[36]) stating that the concept of meme is a less developed Sign. Meme is thus described in memetics as a sign without its triadic nature. Charles Sanders Peirce's semiotic theory involves a triadic structure: a sign (a reference to an object), an object (the thing being referred to), and an interpretant (the interpreting actor of a sign). For Deacon and Kull, the meme is a degenerate sign, which includes only its ability of being copied. Accordingly, in the broadest sense, the objects of copying are memes, whereas the objects of translation and interpretation are signs.

Others have pointed to the fact that memetics reduces genuine social and communicative activity to genetic arguments, and this cannot adequately describe cultural interactions between people. For example, Henry Jenkins, Joshua Green, and Sam Ford, in their book Spreadable Media (2013), criticize Dawkins' idea of the meme, writing that "while the idea of the meme is a compelling one, it may not adequately account for how content circulates through participatory culture." The three authors also criticize other interpretations of memetics, especially those which describe memes as "self-replicating", because they ignore the fact that "culture is a human product and replicates through human agency."[37] In doing so, they align more closely with Shifman's notion of Internet Memetics and her addition of the human agency of stance to describe participatory structure.

Mary Midgley criticizes memetics for at least two reasons:[38]

"One, culture is not best understood by examining its smallest parts, as culture is pattern-like, comparable to an ocean current. Many more factors, historical and others, should be taken into account than only whatever particle culture is built from. Two, if memes are not thoughts (and thus not cognitive phenomena), as Daniel C. Dennett insists in "Darwin's Dangerous Idea", then their ontological status is open to question, and memeticists (who are also reductionists) may be challenged whether memes even exist. Questions can extend to whether the idea of "meme" is itself a meme or is a true concept. Fundamentally, memetics is an attempt to produce knowledge through organic metaphors, which as such is a questionable research approach, as the application of metaphors has the effect of hiding that which does not fit within the realm of the metaphor. Rather than study actual reality, without preconceptions, memetics, as so many of the socio-biological explanations of society, believe that saying that the apple is like an orange is a valid analysis of the apple."[39]

Like other critics, Maria Kronfeldner has criticized memetics for being based on an allegedly inaccurate analogy with the gene; alternately, she claims it is "heuristically trivial", being a mere redescription of what is already known without offering any useful novelty.[40]

New developments

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Alternative definitions

Memetic analysis


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Research methodologies that apply memetics go by many names: Viral marketing, cultural evolution, the history of ideas, social analytics, and more. Many of these applications do not make reference to the literature on memes directly but are built upon the evolutionary lens of idea propagation that treats semantic units of culture as self-replicating and mutating patterns of information that are assumed to be relevant for scientific study. For example, the field of public relations is filled with attempts to introduce new ideas and alter social discourse. One means of doing this is to design a meme and deploy it through various media channels. One historic example of applied memetics is the PR campaign conducted in 1991 as part of the build-up to the first Gulf War in the United States.[54]

The application of memetics to a difficult complex social system problem, environmental sustainability, has recently been attempted at[55] Using meme types and memetic infection in several stock and flow simulation models, Jack Harich has demonstrated several interesting phenomena that are best, and perhaps only, explained by memes. One model, The Dueling Loops of the Political Powerplace,[56] argues that the fundamental reason corruption is the norm in politics is due to an inherent structural advantage of one feedback loop pitted against another. Another model, The Memetic Evolution of Solutions to Difficult Problems,[57] uses memes, the evolutionary algorithm, and the scientific method to show how complex solutions evolve over time and how that process can be improved. The insights gained from these models are being used to engineer memetic solution elements to the sustainability problem.

Another application of memetics in the sustainability space is the crowdfunded Climate Meme Project[58] conducted by Joe Brewer and Balazs Laszlo Karafiath in the spring of 2013. This study was based on a collection of 1000 unique text-based expressions gathered from Twitter, Facebook, and structured interviews with climate activists. The major finding was that the global warming meme is not effective at spreading because it causes emotional duress in the minds of people who learn about it. Five central tensions were revealed in the discourse about [climate change], each of which represents a resonance point through which dialogue can be engaged. The tensions were Harmony/Disharmony (whether or not humans are part of the natural world), Survival/Extinction (envisioning the future as either apocalyptic collapse of civilization or total extinction of the human race), Cooperation/Conflict (regarding whether or not humanity can come together to solve global problems), Momentum/Hesitation (about whether or not we are making progress at the collective scale to address climate change), and Elitism/Heretic (a general sentiment that each side of the debate considers the experts of its opposition to be untrustworthy).[59]

Ben Cullen, in his book Contagious Ideas,[60] brought the idea of the meme into the discipline of archaeology. He coined the term "Cultural Virus Theory", and used it to try to anchor archaeological theory in a neo-Darwinian paradigm. Archaeological memetics could assist the application of the meme concept to material culture in particular.

Francis Heylighen of the Center Leo Apostel for Interdisciplinary Studies has postulated what he calls "memetic selection criteria". These criteria opened the way to a specialized field of applied memetics to find out if these selection criteria could stand the test of quantitative analyses. In 2003 Klaas Chielens carried out these tests in a Masters thesis project on the testability of the selection criteria.

In Selfish Sounds and Linguistic Evolution,[61] Austrian linguist Nikolaus Ritt has attempted to operationalise memetic concepts and use them for the explanation of long term sound changes and change conspiracies in early English. It is argued that a generalised Darwinian framework for handling cultural change can provide explanations where established, speaker centred approaches fail to do so. The book makes comparatively concrete suggestions about the possible material structure of memes, and provides two empirically rich case studies.

Australian academic S.J. Whitty has argued that project management is a memeplex with the language and stories of its practitioners at its core.[62] This radical approach sees a project and its management as an illusion; a human construct about a collection of feelings, expectations, and sensations, which are created, fashioned, and labeled by the human brain. Whitty's approach requires project managers to consider that the reasons for using project management are not consciously driven to maximize profit, and are encouraged to consider project management as naturally occurring, self-serving, evolving process which shapes organizations for its own purpose.

Swedish political scientist Mikael Sandberg argues against "Lamarckian" interpretations of institutional and technological evolution and studies creative innovation of information technologies in governmental and private organizations in Sweden in the 1990s from a memetic perspective.[63] Comparing the effects of active ("Lamarckian") IT strategy versus user–producer interactivity (Darwinian co-evolution), evidence from Swedish organizations shows that co-evolutionary interactivity is almost four times as strong a factor behind IT creativity as the "Lamarckian" IT strategy.


See also


  1. ^ Dawkins, Richard (1981). The selfish gene (Repr. with corr ed.). Oxford: Oxford Univ. Pr. ISBN 978-0-19-857519-1.
  2. ^ Blackmore, Susan J. (2000). The meme machine. Oxford: Oxford university press. ISBN 978-0-19-286212-9.
  3. ^ Dennett, Daniel Clement (1995). Darwin's dangerous idea: evolution and the meanings of life. New York London Toronto [etc.]: Simon and Schuster. ISBN 978-0-684-80290-9.
  4. ^ Dennett, Daniel Clement (2017). From bacteria to Bach and back: the evolution of minds. London: Allen Lane. ISBN 978-0-241-00356-5.
  5. ^ Dennett, Daniel Clement (1995). Darwin's dangerous idea: evolution and the meanings of life. New York London Toronto [etc.]: Simon and Schuster. p. 344. ISBN 978-0-684-80290-9.
  6. ^ Dennett, Daniel Clement (1991). Consciousness explained. Boston: Little, Brown. p. 210. ISBN 978-0-316-18065-8.
  7. ^ Deutsch, David (2012). The beginning of infinity: explanations that transform the world. London: Penguin Books. ISBN 978-0-14-027816-3.
  8. ^ Aunger, Robert, ed. (2003). Darwinizing culture: the status of memetics as a science (Repr ed.). Oxford: Oxford Univ. Press. pp. 163–173. ISBN 978-0-19-263244-9.
  9. ^ Richerson, Peter J.; Boyd, Robert (2008). Not by genes alone: how culture transformed human evolution (Paperback ed., [Nachdr.] ed.). Chicago, Ill.: University of Chicago Press. ISBN 978-0-226-71212-3.
  10. ^ Shifman, Limor (2014). Memes in digital culture. The MIT Press essential knowledge series. Cambridge, Massachusetts: The MIT Press. ISBN 978-0-262-52543-5.
  11. ^ L., Hull, David (2001). Science and selection : essays on biological evolution and the philosophy of science. Cambridge University Press. ISBN 0-521-64339-2. OCLC 876723188.((cite book)): CS1 maint: multiple names: authors list (link)
  12. ^ a b Shermer, Michael (2002-11-14). The Skeptic Encyclopedia of Pseudoscience: [2 Volumes]. ABC-CLIO. ISBN 978-1-57607-653-8.
  13. ^ Hofstadter, Douglas (1996-04-04). Metamagical themas: questing for the ... - Google Books. Basic Books. ISBN 978-0-465-04566-2. Retrieved 2010-02-18.[permanent dead link]
  14. ^ Hofstadter, Douglas (1996-04-04). Metamagical themas: questing for the ... - Google Books. Basic Books. ISBN 978-0-465-04566-2. Retrieved 2010-02-18.[permanent dead link]
  15. ^ "Media Virus!". Rushkoff. Retrieved 14 April 2019.
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  18. ^ "The Journal of Ideas (ISSN 1049-6335): Contents". Retrieved 14 April 2019.
  19. ^ Semon, Richard Wolfgang (1921). The Mneme. London: Allen & Unwin. Retrieved 17 June 2019.
  20. ^ Dawkins, Richard (2006-03-16). The Selfish Gene: 30th Anniversary edition. Oxford UP. p. 182. ISBN 9780191537554.
  21. ^ Hull, David L. (2001-01-04), "Taking memetics seriously: Memetics will be what we make it", Darwinizing CultureThe Status of Memetics as a Science, Oxford University Press, pp. 43–67, doi:10.1093/acprof:oso/9780192632449.003.0003, ISBN 978-0-19-263244-9, retrieved 2022-12-18
  22. ^ Blackmore, Susan (2003). "Consciousness in meme machines". Journal of Consciousness Studies. Imprint Academic.
  23. ^ McNamara, Adam (2011). "Can we Measure Memes?". Frontiers in Evolutionary Neuroscience. 3: 1. doi:10.3389/fnevo.2011.00001. PMC 3118481. PMID 21720531.
  24. ^ Duch, Włodzisław (2021). "Memetics and Neural Models of Conspiracy Theories". Patterns. 2 (11): 100353. doi:10.1016/j.patter.2021.100353. PMC 8600249. PMID 34820645.
  25. ^ Aunger, Robert. "Darwinizing culture: The status of memetics as a science." (2001).
  26. ^ "Journal of Memetics - Evolutionary Models of Information Transmission". Journal of Memetics. Archived from the original on 10 August 2011. Retrieved 17 September 2010.
  27. ^ Dawkins, R. (1982) "Replicators and Vehicles" King's College Sociobiology Group, eds., Current Problems in Sociobiology, Cambridge, Cambridge University Press, pp. 45–64. "A replicator may be defined as any entity in the universe of which copies are made."
  28. ^ Radim Chvaja (2020). "Why Did Memetics Fail? Comparative Case Study". Perspectives on Science. 28 (4): 542–570. doi:10.1162/posc_a_00350.
  29. ^ Cao, Rosa (December 2020). "Crowding out Memetic Explanation". Philosophy of Science. 87 (5): 1160–1171. doi:10.1086/710518. ISSN 0031-8248. S2CID 225622281.
  30. ^ Lankshear, Colin (2011). New literacies everyday practices and social learning. Open University Press. ISBN 978-1-283-26917-9. OCLC 1306561905.
  31. ^ Shifman, Limor; Thelwall, Mike (December 2009). "Assessing global diffusion with Web memetics: The spread and evolution of a popular joke". Journal of the American Society for Information Science and Technology. 60 (12): 2567–2576. doi:10.1002/asi.21185.
  32. ^ Shifman, Limor (2014). Memes in digital culture. Cambridge, Massachusetts. ISBN 978-1-4619-4733-2. OCLC 860711989.((cite book)): CS1 maint: location missing publisher (link)
  33. ^ Lankshear, Colin; Knobel, Michele (2019). "Memes, Macros, Meaning, and Menace: Some Trends in Internet Memes". The Journal of Communication and Media Studies. 4 (4): 43–57. doi:10.18848/2470-9247/cgp/v04i04/43-57. ISSN 2470-9247. S2CID 214369629.
  34. ^ Benitez-Bribiesca, Luis (2001): Memetics: A dangerous idea Archived 2009-07-04 at the Wayback Machine. Interciecia 26: 29–31, p. 29.
  35. ^ Terrence Deacon, The trouble with memes (and what to do about it). The Semiotic Review of Books 10(3).
  36. ^ Kull, Kalevi (2000). "Copy versus translate, meme versus sign: development of biological textuality". European Journal for Semiotic Studies. 12 (1): 101–120.
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