|First appearance||Fantastic Four #2 (January 1962)|
Amazing Adult Fantasy #7 (December 1961, retroactive)
|Created by||Stan Lee (writer)|
Jack Kirby (artist)
|Place of origin||Skrullos; later moved capital to Tarnax IV (destroyed)|
|Pantheon||Kly'bn, Sl'gur't, and Zorr'Kiri|
|Notable members||See Known Skrulls|
|Inherent abilities||Advanced shapeshifting|
The Skrulls are a fictional race of extraterrestrial shapeshifters appearing in American comic books published by Marvel Comics. The Skrulls first appeared in Fantastic Four #2 and were created by Stan Lee and Jack Kirby. They originated from the planet Skrullos and their empire is located in the Andromeda Galaxy. Their infiltration of Earth was a major event in the Marvel Comics universe as shown in the crossover event Secret Invasion.
The Skrulls have made numerous appearances in animated television and video games before making their live-action cinematic debut in the Marvel Cinematic Universe film Captain Marvel (2019). Skrulls have also appeared in Spider-Man: Far From Home (2019) and the Disney+ series WandaVision, What If...? (both 2021) and Secret Invasion (2023), and the film The Marvels (2023).
The Skrulls first appeared in Fantastic Four #2 and were created by Stan Lee and Jack Kirby. The Skrulls next appear in Fantastic Four #18. The four who impersonated the members of the Fantastic Four next appear in The Avengers #92. The concept of the Skrulls would be explored and used throughout multiple storylines, spin-off comics, and derivative projects.
Millions of years ago, in the main Marvel continuity, the Celestials perform genetic experiments on the reptilian ancestors of the Skrulls, resulting in three branches of Skrulls: the Prime, the Deviants and the Eternals. Eventually, the three branches battled with one another and the Deviant branch – due to possessing the innate ability to shapeshift – were triumphant, and wiped out all members of the other two races until only two were left: the Skrull Eternal Kly'bn, and the Prime Skrull of the original non-Deviant Skrull race. Kly'bn implored them to spare his life, as killing him would kill part of their heritage. The Deviants' leader Sl'gur't eventually fell in love with Kly'bn, with the two eventually becoming the gods of the Skrull pantheon. Taking the name the Prime Skrull, the last original Skrull escaped to Earth in the 20th century and later became a member of the Underground Legion. From that point on, the Skrulls began to expand their territory. The Deviant branch later split into two more groups, the modern Skrulls and an anomaly called the Dire Wraiths.
The Skrulls originated from the planet Skrullos, and were originally a mercantile civilization, primarily interested in free trade and willing to share their technology with all races they deemed worthy. When they encountered a new race, they simply transformed themselves to resemble that race. The Skrull Empire that resulted from these contacts was based on free trade and mutual cooperation.
Main article: Kree–Skrull War
Eventually, the Skrulls developed long-distance space travel; a great tour of the universe was undertaken, led by Emperor Dorrek. Finally, the Skrull delegation reached the planet Hala, home to the then-barbaric Kree and the peaceful Cotati, and held a contest to determine which of the races would represent Hala to the Skrull Empire. Seventeen members of each race were taken to different uninhabited planetoids, where they were left with sufficient supplies for one year. At the end of that period, whichever group had done the most with themselves would be adjudged the most worthy. The Kree were taken to Earth's Moon where they built a great city, while the Cotati were taken to another barren world in a different solar system and used their abilities to create a beautiful park. Realizing that the Cotati were going to win, the enraged Kree killed all of them. When the Kree revealed that they had solved the question of who would represent Hala to the Skrulls by destroying their opposition, the Skrull delegates were appalled and vowed that Hala would forever be banned from their circle of favored worlds—so the Kree massacred them as well, took over the Skrull landing spaceship by force, and developed their own technology from it.
Because of the immense distances involved, decades passed before the Skrulls learned of the Kree's activities. By this time, however, it was too late. The Kree were now advanced and audacious enough to attack the Skrulls in their home galaxy. During the millennia that followed, the Kree aggression forced the Skrulls to become a militaristic civilization, and the Skrulls eventually developed the vicious streak needed to conduct intergalactic war. Their entire culture was remade in the warrior image. The Kree–Skrull War continued for thousands of years with only brief interruptions. The Skrull Throneworld is later moved from Skrullos to the more central Tarnax IV, with their empire encompassing over fifty worlds. The Skrulls also created the first Cosmic Cube, which later became the Shaper of Worlds; the Shaper related that, upon attaining self-awareness, he lashed out blindly, destroying two-thirds of the Skrull Empire and casting the civilization back into barbarism (suggesting this occurred before the war with the Kree broke out).
The Kree Accuser Ronan had wrested control of the Kree Empire from the Supreme Intelligence and attacked the Skrulls, reigniting hostilities. The Avengers become involved once the Super-Skrull kidnapped Captain Marvel and, in turn, they battled a Kree Sentry robot, three of the four original Skrulls that had fought the Fantastic Four, S.H.I.E.L.D. agents, rogue Inhumans and legions of Skrulls. The Supreme Intelligence ended the immediate conflict by revealing to the Avengers' ally Rick Jones that he had vast mental potential, which was then used to freeze all combatants in place. The Supreme Intelligence announced that, while the Kree and Skrulls had reached genetic dead ends, the human race displayed incredible untapped potential. It is revealed years later that the conflict produced a Kree-Skrull hybrid, the Hulkling, who was the child of one-time lovers Captain Marvel and the Skrull Princess Anelle.
The Skrulls became aware of Earth when they discovered a space warp linking Earth and Throneworld. Later, they discovered Earth possesses an equidistant link to Hala. They see Earth as a world to conquer, but they are content to hold off a full invasion. Because of their caution, the Skrulls have sent only a handful of scout warriors to infiltrate Earth, but failed to take into account the presence of the modern superhero.
In 1958, Skrulls attempted to sabotage Earth's space program. They battled the 3-D Man, and set the Cold Warrior against the 3-D Man. Later, a small scouting party used their shape-shifting powers to impersonate the Fantastic Four, committing crimes so the country would turn against the Fantastic Four, the only ones with the power to stop them at the time. But the Fantastic Four were able to locate the Skrulls and tricked their leaders into believing the Earth was full of threats. The fleet left and Mr. Fantastic made three of the four Skrulls that were left behind shape-shift into cows and he hypnotized them to remember nothing about their true heritage.
In retaliation, the Skrull Emperor Dorrek VII dispatched Kl'rt, a Skrull who was made into the Super-Skrull, to Earth to defeat the Fantastic Four. Kl'rt possessed the combined powers of the entire Fantastic Four (in addition to shape-shifting and hypnotism), and he held the team at bay until Mister Fantastic discovered the source of his power and used a miniature device to jam the ray, before the Human Torch imprisoned him inside a crater. The Super-Skrull posed as the jailed Dr. Franklin Storm, after transporting him to the Skrull Throneworld, and battled the Fantastic Four as the Invincible Man. Another Skrull, ultimately caused the death of Dr. Storm by attaching a bomb to his chest before he was transported back, but Storm turned to the floor, saving the Fantastic Four. The Fantastic Four later traveled to Tarnax IV and found the Skrull responsible for the murder of Susan and Johnny Storm's father (which turned out to be the Skrull Warlord Morrat). Skrulls were involved in other events, such as sending the Super-Skrull to battle Captain Mar-Vell, and abducting the Thing as a contestant in the Skrull Games.
Later, the cosmic being Uatu the Watcher adjudicates a duel between champions from both races: Colonel Bel-Dann of the Kree and Warlord Raksor of the Skrulls. After one year of conflict, the duel remained deadlocked and was ultimately abandoned, with no resolution. Years later, during a conflict with Xandar that included the Fantastic Four and the Champions of Xandar, Emperor Dorrek VII's wife Empress R'klll staged a coup and killed him to become ruling Empress.
Empress R'klll's reign was short-lived, however, as the greatest blow to the Skrulls arrived in the form of the World Devourer Galactus. After his Herald Nova annihilated the Skrull fleet, Galactus devoured Tarnax IV, killing 7 billion Skrulls, including R'klll and Princess Anelle. With the destruction of the central government, the Skrull Empire collapsed into hundreds of bickering factions. Their galaxy was filled with Governors who declared themselves to be Emperor of the Skrulls and soon, a civil war began. The Shi'ar maintained a heavy presence in the former Empire as well, constantly dealing with pockets of resistance.
Later, Reed Richards discovered that yet another Governor-turned-Emperor, Zabyk, had created a genetics bomb, which, when it exploded, made all the Skrulls lose their shape-shifting abilities. Whatever form they were in, they were stuck in at the time of the explosion.
In desperation, since without their shape-shifting powers they were vulnerable, they managed to kidnap the herald of Galactus Nova in an attempt to get Galactus to destroy Kree worlds for them. Their plot failed when the Silver Surfer helped Galactus rescue Nova in exchange for his freedom from Earth.
When the Celestials made an appearance, this scared the Skrulls into starting a second war with the Kree to show they were not powerless. The attack was first made by Kylor, one of five governors claiming to be Emperor. He had a spy in the Kree Empire, Nenora, who gave him the early edge. They tried to hide their secret weakness at all costs from the Kree.
Kylor was eventually betrayed by Nenora, as was the entire Skrull race. She assumed power of the Empire and wished to keep it for herself. It was Empress S'byll who stepped up to control the Skrulls and finish the war. She was able to resurrect the Super-Skrull and he returned to find the empire in shambles. They tricked the Silver Surfer into attacking a Badoon fleet, which he was not aware were allied with the Kree. Since the Kree waged war on the Surfer, he allied himself with the Skrulls to save Zenn-La from destruction.
To restore their powers, the Super-Skrull's DNA needed to be transferred to S'byll, because only a female could spread the ability to the other Skrulls. The Surfer helped them to power a machine that would return the Empress' shape-shifting abilities. The device was successful. With this power, she was able to restore every Skrull she touched and became the true Empress of the Skrulls.
She and the Super-Skrull waged war with the Kree with the help of the Silver Surfer. She replaced the Super-Skrull with Captain Reptyl and he helped to change her into an emotionless reptile. But Reptyl was betrayed by one of his own and the Skrull fleet was utterly destroyed by the Kree. S'byll and the Silver Surfer went alone to attack the Kree homeworld of Hala.
Gaining the knowledge that Nenora was actually a Skrull gave them the advantage. S'byll gave her shape-shifting abilities back to her, crippling her and showing the Kree her true self. A peace treaty was created.
The Skrulls eventually broke that peace treaty, as they kept their military drive and prefer to fight rather than negotiate. Among the other races contained within the Skrull Empire are the Druff, Guna, Kallusina, Morani, Pheragot, Queega, Tekton, and Yirbek races. Other races remain independent, whether through treaties or open resistance. Such races include the Clegrimites, Gegku, Krylorians, Wilameanis, and Xandarians. The Skrull-Xandarian war with the Xandarians recently ended with the destruction of Xandar by the supervillainess Nebula.
Skrulls are, meanwhile, involved in numerous other activities. They were revealed to have conquered a medieval planet and abandoned it after its inhabitants perished. Skrull Prime Ten battled the Fantastic Four and Captain Mar-Vell. The Skrulls sent their own representative to witness the fate of the Phoenix, at her trial by the Shi'ar. There is a lone appearance of Queen Akilll of the Skrulls. A lone Skrull poisoned Vera Gantor to force the Avengers to seek the Resurrection Stone on his behalf. The Skrulls sent General Zedrao to give the dying Captain Mar-Vell the Royal Skrull Medal of Honor. The Skrulls had captured a Kree spy who had infiltrated into the Skrulls by using a symbiote.
A group of genetically-bred Skrulls called the Warskrulls began infiltrating the Shi'ar Empire. They used a Nexus Amplifier to assume total physical discorporation of their targets, adopting their superpowers along with their physical appearance. They captured and replaced the Starjammers with members of their race, as well as key components in government. The Warskrulls, acting under the direction of the paramount among them, posing as a Shi'ar called the Chamberlain, began committing murders and even genocide upon various Shi'ar affiliated races, such as the P!ndyr, in the name of the Shi'ar Empire. One Warskrull replaced the Imperial Consort, Professor X, called himself Warlord, and took mental control over Lilandra, the Majestrix of the Shi'ar. They were eventually defeated by the combined forces of Deathbird and the X-Men and later the Shi'ar vowed to weed out any other Warskrulls.
Some time later, a Skrull called Paibok the Power Skrull mounted a campaign to invade Earth, using another Skrull called Lyja as an infiltrator. The plan involved secretly replacing the real Alicia Masters, the Thing's blind lover. The plan fell through when she instead seduced the Human Torch and the two eventually married one another. The long-running facade eventually came to light and the group discovered that the real, human Alicia was actually being held captive by the Skrull nation and Lyja defected, having fallen in love with the Human Torch. The Fantastic Four rescued her, but Lyja seemingly died in the process as she sacrificed herself to save Johnny when Paibok attacked.
Paibok later revealed to Devos the Devastator that she was still alive. Paibok, with the assistance of Devos, awoke Lyja from her comatose state and gave her the power to fly and fire energy blasts from her hands (she called herself "Lyja the Lazerfist"). These powers were later revealed to come from a special device implanted in her. Joining them in search of vengeance, the three arrived on Earth, and battled the Torch at Empire State University. The Torch panicked, used his nova flame and destroyed the campus. Alongside Devos and Paibok, Lyja witnessed a battle between the Fantastic Four and an alternate Fantastic Four. However, Lyja still bore feelings for the Torch and once again betrayed her cohorts to side with the Fantastic Four. This ended with the Fantastic Four destroying the "Skrull War World", the Skrulls' largest attack ship.
A group of Skrulls traveled to Earth, where they began plotting to invade the planet once again. They were, however, at some point approached by Apocalypse, who wanted to lure together 12 powerful mutants that he needed to ascend to godhood. At the same time, the body of the Living Monolith, who had been hurled out into the far reaches of space by Thor, is found and brought back to Earth. On Earth, a mutant Skrull named Fiz snuck into Xavier's mansion to tell them urgent news of trouble: the Skrulls have allied themselves with someone powerful and the only thing he knows is that there are 12 mutants that cannot be harmed and that something terrible is going to happen. Fiz also reveals the existence of a subtype of Skrulls that possess innate powers like Earth's mutants. Deemed outcasts by the other Skrulls, they were hunted by the Empire.
After the eventual defeat of Apocalypse, Professor X went into space to train them, giving them the name Cadre K. Cadre K and Xavier returned to Earth during the Ruul's attempt to overtake the planet by turning it into an intergalactic prison. A wounded Z'Cann attempted to pass on vital information to Rogue. Their physical contact affected Rogue's powers for some time. The information was received and the X-Men were reunited with Bishop.
Main article: Annihilation (comics)
The Skrull Empire is the first of the major interstellar empires to be invaded by the forces of Annihilus. The Annihilation Wave's superweapon, the Harvester of Sorrow, physically destroys dozens of Skrull planets. In the aftermath, the Skrulls unsuccessfully attempt to convince the Hulkling to become their new Emperor.
A group of Warskrulls attacked a remote Shi'ar space station and took it over. Disguised as Shi'ar, they were taken aboard the space station willingly and slaughtered all those aboard, except a few needed to keep working the ship. The group, however was eventually defeated by a group of X-Men (Darwin, Havok, Marvel Girl, Nightcrawler, Professor X and Warpath). These Warskrulls are, however, revealed to be cyborgs.
In the Secret Invasion series by Brian Michael Bendis and Leinil Yu, a flailing Skrull Empire, under the leadership of newcomer Queen Veranke, made an all-out effort to infiltrate and conquer Earth. It was revealed that the Skrulls had infiltrated Earth for quite some time by capturing and replacing many of Earth's heroes. They followed this infiltration with their main assault group. This wave of Skrulls then formed new versions of the Super-Skrull, with each possessing the abilities of several different heroes. Their invasion ultimately failed, costing the life of Veranke, and reducing their numbers even further. Even their gods were slain, and the Japanese God of Evil Amatsu-Mikaboshi took over their Pantheon, leading to the events of Chaos War.
After the events of Secret Invasion, the Skrulls, now reduced in numbers, became very segmented with various factions rising up around Warlords that began plotting to rise back to power by orchestrating events for their own benefit.
During the Infinity storyline, several planets that the Skrulls settled on began being targeted by the Builders and while the Silver Surfer tried to save most of them, the Skrulls' population was once more decimated. Kl'rt the Super-Skrull later represented the Skrull Empire as a member of the Galactic Council. After the Avengers' victory over the Builders and Thanos, Kl'rt becomes the new Emperor of the Skrulls, where they settle on Tarnax II.
Meanwhile, Cadre K had settled on a ranch in South Dakota, while another group of Skrulls remain undercover on Earth as the "Warner" family, who are gathering data for the next Skrull invasion. The Warner family participate in day-to-day life, but end up stranded on Earth, completely cut off from the Empire.
During the search of the Infinity Stones in the Infinity Countdown storyline, Razor reveals the true origin of the Tree of Shadows; according to Razor, the Tree of Shadows was created eons ago by one of the Elders of the Universe known as the Gardener, who tried to bring life to the Null Space and planted it in the Darkforce Dimension. Eventually the Null Space and the Tree of Shadows were discovered by a more prehistoric version of the Shi'ar and the Skrulls, a find that erupted in a war between the two alien races. The war raged until one member of the Shi'ar was able to grasp one of the seeds of the Tree. His physiology, fear and beliefs shaped the birth of the most vicious killing machine the universe had ever seen, the first Raptor and while the Skrulls, being shape-shifters, were still able to hold out against the Shi'ar, they eventually were nevertheless slaughtered by Raptor Prime, a creature of instinct and adaptation which quickly learnt to use their enemies' shape-shifting abilities as well.
In the "Road to Empyre," Teddy Altman is made a mysterious offer at the cost of leaving Billy Kaplan. He accepts the offer which was to become the new ruler of the Kree-Skrull Alliance, adopting the mantle of "Dorrek VIII," and beginning the preparations of invading the Earth for "the final war."
General G'iah, under the alias of "Gloria Warner", sends an encrypted message to the Skrull High Command about the progress of Project Blossom. Her mate, Colonel Klrr, uses the alias of "Carl Warner" and they have three daughters named Alice, Ivy, and Madison. After losing her husband to their handler and rescuing Ivy, General G'iah obtains a sample of a Cotati from a lab while explaining the history of the Kree and the Skrull's feud. At their motel, they are ambushed by a Kree operative. General G'iah leads her daughters to track the assailant to a Kree family. After G'iah's daughters prevent her from retaliating against them, the patriarch of the Kree family that blew up their motel room suddenly receives a message on his Kree-tech cellphone by Dorrek VIII, stating to all Kree and Skrull soldiers in the field that the Kree and Skrull armadas have united to face a common enemy. They are coming to Earth to destroy them as he speaks. The enemy is eventually revealed to be the Cotati, who are returning, as well as the Celestial Messiah. In the Kree/Skrull Alliance, Dorekk VIII has gained Captain Glory, Mur-G'nn, Tanalth the Pursuer, and the Super-Skrull as his inner circle. When the Avengers learned about the Cotati's true motives, they decided to work together to fight the Cotati and prevent them from eliminating all animal life. As Dorekk VII allows the Black Panther to wield the Sword of Space, Captain Glory has a private chat with Tanalth the Pursuer and his scans reveal that she is not a Kree. It is revealed that Tanalth the Pursuer is actually Empress R'Klll in disguise, having survived the destruction of Tarnax IV. When Captain Glory states that her grandson spent too much time with the humans, R'Klll notes that it brought out the Hulking's heroism and states that they will have to do something about that.
Skrulls are green-skinned reptilian humanoids with large pointed ears, red or green eyes, and chins with several vertical folds in the skin below their mouths. Skrulls are known for genetic and molecular instability and genetic diversity, due to Celestial experimentation creating the Skrull Deviants. The Skrulls are known for their physical malleability and ability to shapeshift to any size, shape, or color at will, taking on the appearance but not the characteristics of other beings and objects. Skrulls are able to assume virtually any form, be it organic (e.g., cows ) or inorganic (e.g., lamps ). As a result, the Skrulls excel at spying and infiltration. Skrulls are also able to use their shapeshifting abilities to form weapons (e.g., blades and clubs) with parts of their bodies, making them dangerous hand-to-hand combatants. Their sexual dimorphism is roughly the same of a human being, with the exception that the males are mostly bald and have no facial hair, while females have full head of greenish-black hair, but their sexual orientation is more complex, since they can change their genders at will. Indeed, Xavin once casually said that, for a Skrull, changing gender is comparable to a human being changing their hair color. As for their reproductive system, just like the other reptilians, the females do not give live birth, but instead lay eggs and the newborns are known as "hatchlings". However, despite being reptilian, the females also possesses hair and mammary glands to nurture their newborn, just like a human being.
The Skrulls later developed the ability to render themselves undetectable when using their shapeshifting abilities, even from telepaths and those with superior senses. Reed Richards discovered a method to use technology to negate this ability and reveal a disguised Skrull's true form.
Courtesy of their advanced technology, the Skrulls have also been able to augment their abilities in certain warriors, such as the Super-Skrull, Power Skrull and the Warskrulls—all elite groups who, with special encoding, are able to emulate the powers as well as the appearance of their biophysical templates or are bestowed entirely unique supertropical abilities all their own (in the case of Paibok).
The Skrulls have been shown to suffer from various diseases. Some Skrulls are born with a defect that makes them unable to shapeshift, however, this defect also enhances their strength to those Skrulls including Titannus, Talos and Raze. The Skrulls can also suffer from cancer, as shown with Kl'rt the Super-Skrull and Prince Klundirk. In an alternate universe, Skyppi suffered from a form of tuberculosis which was common to the Skrull species.
Just like the Kree, the Skrulls were evolutionarily stagnant, unable to evolve. However, over the millennia Skrull mutants have been born with the K-gene as opposed to the Human Mutants X-Gene, meaning that now this might not be the case anymore.
After a Skrull dies, they revert to their original form.
The Skrull species speak a variety of languages and dialects, including: Skrullos, Skrullian, the Irdu dialect, the Tligi dialect and the Frti dialect. While the Prime Skrulls had their own language, however, it was forgotten as their species was extinct.
Since the Skrulls are a warrior race, it is hard for them to grasp almost any kind of representational art.
The main Skrull religion is the "Dard'van Sect", their "True Faith", which worships Sl'gur't and Kly'bn. It is unknown about the relation between Zorr'Kiri and the actual pantheon.
In Deadpool vs. Thanos #3, it is revealed that the Skrull equivalent of Hell is called the Shapeless Wastes.
In Amalgam Comics, the Skrull are the native inhabitants of Mars (having amalgamated DC's Green Martians with the Skrulls). Mister X (the Martian Manhunter and Professor X) is an example of a Martian Skrull.
In the alternate universe limited series Earth X, Thanos dwelled in the Realm of the Dead with the entity Death. It is revealed his mother was a Skrull and Death used her secret to make him believe that Death was his mother.
In the one-shot issue Hulk: The End, an alien Recorder robot tells an elderly Bruce Banner, the last human on the planet, that the death of the human race was widely celebrated throughout the universe, so much so that the Kree and the Skrulls had settled their differences in celebration.
The Skrullian Skymaster (real name Skymax) is a founding member of the Squadron Supreme and possesses abilities matching the Super-Skrull from Earth-616. The character first appeared in flashback in Squadron Supreme #1 (1989), and appeared in more detail in Squadron Supreme: New World Order.
On Earth-712, the Skrull Empire has been at war with the Kree Empire. In an attempt to end the bloodshed, the Skrull Empire created a means to imbue their soldiers with extraordinary powers to vanquish the Kree. Skymax was the first and the only soldier subjected to the experiments, because the World Devourer destroyed the Skrull homeworld. Sk'ym'x was in the vicinity of Earth when his starship malfunctioned. He was rescued by American astronaut Joe Ledger (who would later become Doctor Spectrum). Grateful for his rescue, Skymax gave Ledger a Power Prism, which Ledger used to become the costumed adventurer Doctor Spectrum, while Skymax became the costumed adventurer known as the Skrull. Skymax and Ledger both became founding members of the Squadron Supreme, and Skymax remained with the team until member Tom Thumb repaired his ship, allowing him to return to space. Skymax realized that the damage to his ship was beyond repair despite Tom Thumb's efforts, and he chose to remain on Earth in secret, searching for any possibility of other Skrull survivors. Not finding any, Skymax concluded that he was indeed the last of his race; he remained on Earth.
After the Squadron Supreme returned from their adventures on Earth-616, the sorcerer Mysterium sent the surviving members to retrieve others to help in the rebellion against the Global Directorate. Doctor Spectrum was sent after "the Watcher" and discovered Skymax. Renaming himself the Skrullian Skymaster and revealing his true origin to Doctor Spectrum, Skymax rejoined the Squadron Supreme and settled with them on Utopia Isle.
In the 2021 "Heroes Reborn" reality, the Skrullian Skymaster was established as a former member of the Squadron Supreme of America. While trying to break up the protests caused by mutants over a bill that would start the Mutant Registration Act, the Skrullian Skymaster killed Mystique with his flame attacks. This caused Rogue to use her life-draining powers to avenge her adoptive mother by killing the Skrullian Skymaster.
In the Ultimate Marvel reality, Skrulls are led by the billion-year-old Skrull Emperor Kl'rt, as their off shoot Chitauri exist whom they call terrorists. They have extremely advanced technology, but have not been observed to shapeshift. These Skrulls were seen only in an alternate timeline in which Reed Richards contacted their world via his teleporter; the events leading to that timeline were altered in Ultimate Fantastic Four #29 so that contact was never made.
In the MC2 universe, Torus Storm, the son of Lyja and Johnny Storm, is a Skrull/Human hybrid, who inherited the powers of both his parents.
In an alternate timeline set 30 years into the future, the Skrull Empire was united with the Kree Empire under the rulership of Emperor Dorrek VIII and they were successful in eliminating all life on Earth.
On Earth-10021, the Skrulls were victorious in the end of Secret Invasion and the heroes were defeated. In this reality, the Skrulls were coexisting along with the humans on Earth and were seen by humans as people with good intentions. Despite helping the humans in medical care and technology, in reality, they are secretly turning the humans into Skrulls like themselves, including Norman Osborn, who had married Queen Veranke, while the heroes had been hiding in Wakanda, since now they were seen as criminals. In the end, the heroes spread a virus which killed every Skrull and the humans who had been turned into Skrulls, except for Osborn, who had been disguised as Iron Man. Osborn revealed that he had orchestrated the plan all along, but he was killed by Bucky Barnes. The heroes were then taken by the U.S. government as world criminals.
Actor Tye Sheridan revealed in a 2019 interview that the Skrulls were originally intended to be the main antagonists in an earlier cut of Dark Phoenix before they were replaced by the D'Bari amidst re-shoots.
The Skrulls appear in media set in the Marvel Cinematic Universe: